Family Pieridae  Duponchel, [1835] ( [1] )

Subfamily Dismorphiinae  Schatz, 1886

Tribus Leptideini  Verity, 1947

 

Enumeratio insectorum musei Bilbergii: 76. TS: Papilio sinapis Linnaeus, 1758.

 

amurensis Ménétriès, 1859  (Leucophasia). Bulletin de la Academie impériale des Sciences, Saint-Pétersbourg, Ser. 2, 17: 213. TL: “les montagnes de Khingan jusqu’à Pakhale”. Had been originally described as a good species.

·       amurensis Ménétriès, 1859 (Leucophasia). D: Ussuri district; Amur district.

·       emisinapis Verity, [1911]  (Leptidea). Rhopalocera palaearctica, Firenze , [1]: 345. TL: Irkutzk. Had been originally described as a subspecies of amurensis. D: Altai; Sayan; Praebaikal.

·       jacutia P.Gorbunov & Korshunov, 1995  (Leptidea). Dnevnye babochki asiatskoj chasti Rossii, Ekaterinburg: 56. TL: “Якутск, ботсад” [ Yakutsk , botanic gardin]. Had been originally described as a subspecies of amurensis. D: Central Yakutia .

·       vibilioides Verity, [1911]  (Leptidea). Rhopalocera palaearctica, Firenze , [1]: 345. TL: Amur district, Blagovestshensk. Had been originally described as a subspecies of amurensis. D: Transbaikal; environs of Blagovestshensk, Sakhalin .

 

duponcheli Staudinger, 1871  (Leptidia). Catalog der Lepidopteren des europaeischen Faunengebietes, Dresden (ed. 2): 5. TL: “Gal. m.; Ped. Bith. Pont.” S France . Had been originally described as a good species.

·       majae Alberti, 1969  (Leptidea). Deutsche entomologische Zeitschrift, 16: 191; Taf. 1, Fign. 1, 2. TL: “Kislowodsk”. Had been originally described as a subspecies of duponcheli. D: Crimea; Caucasus ; Transcaucasus.

 

litania Churkin, 2004 (Leptidea). Helios, Moscow , 5: 138 – 140; figs. 1 a , b, e, g; pl. 10, figs. 1 – 4. TL: “Tadjikistan, Peter the Great Range , 23 km SE Tadzhikobad [sic!], Ganishou v.”. Had been originally described as a good species.

·       litania Churkin, 2004 (Leptidea). D: TL.

 

morsei Fenton, 1881  (Leptosia). Proceedings of the Zoological Society of London , 1881: 855. TL: “Yesso”, Japan . Had been originally described as a good species.

·       major Grund, 1905  (Leptosia). Entomologische Zeitschrift, Frankfurt am Main, 19: 145; Fign. 3, 4. TL: “Kroatien, Hrvatska”. Had been originally described as a subspecies of morsei. D: SW part of ex-USSR’s Europa.

·       micromorsei Veriy, 1947 (Lepidea). Le farfalle diurne de l’Italia, Firenze , 3: 120. TL: Amur region, Pokrovka. Had been originally described as a race of morsei. D: Amur region.

·       morsei Fenton, 1881 (Leptosia). D: Asiatic part of Russia from Baikal to Kuriles up to northern taiga northwards.

·       morseides Verity, [1911]  (Leptidea). Rhopalocera palaearctica, Firenze , [1]: 244. TL: “Sajan: Tonkoun” (Kudrna, 1983: 58). Had been originally described in quadrinominal combination Leptidea sinapis morsei morseides. D: South Ural; Altai; Sayan; Praebaikal.

·       ssp ?  D: Central part of European Russia.

 

reali Reissnger, 1990 (Lepidea). Atalanta, Würzburg, 20: 149 [nom. nov. pro Leptidea lorcovicii Real, 1988, nec Lepidea duponcheli lorkovici Pfeiffer, 1932]. TL: East Pyrenees , 1300 m . Had been originally described as a good species.

·       melanogina Lorkovič, 1993 (Leptidea). Natura Croatia , 2 (1): 21, figs. 25-28. TL: “ Zagreb , Kanal”. Had been originally described as a subspecies of reali. D: Caucasus ( Sochi and Krasnodar ), Bryansk region, Souh and Middle Ural.

·       churkini Korb, in litt. Issyk-Kul lake coasts.

·       yakovlevi Mazel, 2001 (Leptidea). Linneana Belgica, Tervuren, 18 (2): 103 – 104. TL: “ Novosibirsk reg., Berdsk”. Had been originally described as a subspecies of reali. D: . Regions of Tyumen , Tomsk and Novosibirsk .

 

sinapis Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 468. TL: Sweden . Had been originally described as a good species.

·       melanoinspersa Verity, [1911]  (Leptidea). Rhopalocera palaearctica, Firenze , [1]: 344. TL: Turkestan . Had been originally described in quadrinominal combination Leptidea sinapis sinapis melanoinspersa. D: Tian Shan; Darvaz; Alai; West Pamir ; Altai.

·       sinapis Linnaeus, 1758  (Papilio). D: European part; Caucasus; Transcaucasus; Crimea .

Subfamily Coliadinae  Swainson, 1827

Tribus Coliadini  Swainson, 1827

Magazin für Insektenkunde, 6: 284. TS: Papilio hyale Linnaeus, 1758.

 

Subgenus Colias sensu stricto

 

alfacariensis Ribbe, 1905  (Colias). Societas Entomologica, 20: 137. TL: “Sierra de Alfacar”. Had been originally described as an aberratio of hyale.

·       fontainei Reissinger, 1989  (Colias). Neue entomologische Nachrichten, Marktleuthen, 26: 45. TL: Borzhom. Had been originally described as a subspecies of alfacariensis. D: Caucasus ; Transcaucasus; Kopet-Dagh.

·       remota Reissinger, 1989  (Colias). Neue entomologische Nachrichten, Marktleuthen, 26: 146. TL: “Wolsk”. Had been originally described as a subspecies of alfacariensis. D: South of European part from Praecaucasus to line Belgorod – Samara – Orenburg ; South Ural; Crimea .

·       saissanica Reissinger, 1989  (Colias). Neue entomologische Nachrichten, Marktleuthen, 26: 138. TL: “Saisan”. Had been originally described as a subspecies of alfacariensis. D: South-West Altai; Tarbagatai; ?Dzhungaria.

·       vihorlatensis Reissinger, 1989  (Colias). Neue entomologische Nachrichten, Marktleuthen, 26: 112. TL: “Vihorlat-Gebirge”. Had been originally described as a subspecies of australis. D: environs of Vihorlat Mts. in west, south and east of Ukrainian Carpathians (REISSINGER, 1989: 165).

Remark. Y.P.Korshunov (1996: 19) wrote: “Среди бабочек из лесостепи Ю.Урала, З.Сибири, до Енисея, иногда попадаются экземпляры, близкие к южно-европейскому виду Colias sareptensis Staudinger, 1871  (= alfacariensis Ribbe, 1905, australis Verity, 1911).  Здесь можно бы поставить stat.n., если бы ещё в 1876 г . в трудах РЭО, т. 8: 153 – 154 С.Алфераки не вывел этот таксон из обозначения гибрида hyale и erate. С.Альфераки [sic!] представляет sareptensis  “постоянным видоизменением, свойственным южной и юго-восточной России”. Он пишет, что sareptensis крупнее, чем типичные самцы hyale, цвет их крыльев гораздо желтее, часто одинаковый с цветом самцов erate; черная кайма передних и задних крыльев гораздо шире, чем у C. hyale. Самки гораздо больше, чем самцы; цвет крыльев у них белый с небольшой примесью желтоватого; черная кайма передних крыльев доходит до внутреннего угла их, как это у самок erate ab. pallida Stgr. Крылья самок sareptensis гораздо удлиненнее, чем у hyale. Наконец, бабочки летают в тех же местах и в те же сроки, что и hyale и erate. Остается сожалеть, что западные авторы, потратив так много бумаги на обоснование australis, а позже alfacariensis, не обращали внимания на российские публикации”. [In the butterfly collections from forest-steppes of South Ural and West Siberia to Enisej have rare specimens which are close to the south-european species Colias sareptensis Staudinger, 1871 (= alfacariensis Ribbe, 1905; australis Verity, [1911]). I can marked here a  status nova but in 1876 (Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 8: 153 – 154) S.Alpheraky “made” this taxon as a hybrid hyale X erate. S.Alpheraky think that sareptensis is “a constant aberratio from South and South-East Europa”. He wrote that males of sareptensis less than males of hyale, that colour of wings of sareptensis yellower often as in males of erate; black border of wings wider as in C. hyale. Females less than males; wing colour of females white with little yellowish tinge; black border in forewing up to anal border as in females of sareptensis longer than in hyale. Endly butterflies fly syntop and synchron. I’m sorry that the western authors used so many paper to the investigations in case alfacariensis – australis but not used publications of Russian authors]. So the conspecificity of alfacariensis and australis was not good maded, I use here the name alfacariensis.

 

alta Staudinger, 1886  (Colias). Stettiner entomologische Zeitschrift, 47: 200 - 201. TL: “Alai-Gebirge…, Kara Kasuk (Paß) 10,000’ hoch”. Had been originally described as a variety of hyale.

·       alta Staudinger, 1886 (Colias). D: Mountains of Pamir, Gissar, Darvaz and Alai .

·       grieshuberi  Korb, 2004  (Colias).  Alexanor, Paris , 23 (2): 115; Abb. 1. TL: “Barskaun-Schlucht, 10 km südlich von Barskaun, Kirgisien”. Had been originally described as a subspecies of alta. D: Mountains of Tian Shan .

 

hyale Linnaeus, 1758  (Colias). Systema Naturae... (ed. 10), 1: 469. TL: South England . Had been originally described as a good species.

·       altaica Verity, [1911]  (Colias). Rhopalocera palaearctica, Firenze , [1]: 348. TL: “Altai”. Had been originally described as a race of hyale. D: Altai; Tarbagatai; Dzhungaria; North Tian-Shan: Transili Alatau, Kungey Ala-Too, Ketmen and Kirghiz mountain ridges.

·       hyale Linnaeus, 1758 (Colias). D: European part; Crimea; Caucasus ; Transcaucasus.

·       irkutskana Stauder, 1924  (Colias). Mitteilungen der Münchener entomologischen Gesellschaft, 14: 62. TL: Irkutzk. Had been originally described as a subspecies of hyale. D: Sayan; Praebaikal; Transbaikal; Amur district.

·       pallidis Fruhstorfer in Seitz, 1910  (Colias). Die Groβschmetterlinge der Erde. Die Groβschmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart , 9: 164. TL: “Wilui”. Had been originally described as a subspecies of hyale. D: Yakutia; East Siberia .

Subgenus Asiocolias  Korb, 2005 .

TS: Colias christophi Grum-Grshimailo, 1885.

 

 

christophi Grum-Grshimailo, 1885  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 2: 220. TL: “Ak-Su, Artscha-Bulak, Kara-Kasuk, Artscha-Basch-Platze”. Had been originally described as a good species.

·       christophi Grum-Grshimailo, 1885 (Colias). D: Alai and Transalai mountain ridges.

·       kali Korb, 1999 (Colias).  Alexanor, Paris , 20 (8): 452 – 453; fig. 5. TL: “chaîne montagneuse du Turkestan [Turkěstanskij hrěbět]”. Had been originally described as a subspecies of christophi. D: Gissar, Darvaz.

 

Subgenus Eucolias Berger, 1986  

Lambillionea, Bruxelles, Suppl. 86: 7. TL: Colias nastes Boisduval, 1832.

 

alpherakii Staudinger  in Staudinger & Bang-Haas, 1882 (Colias). Berliner entomologische Zeitschrift, 26: 164. TL: “auf dem Alai Gebirge (Fergana Turkestan),... auf den Hazret-Sultan Alpen (Prov. Samarkand)”. Had been originally described as a good species.

·       alpherakii Staudinger in Staudinger & Bang-Haas, 1882 (Colias). D: Darvaz; Vanchsky Mts. in West Pamir; Alai and Transalai mountain ridges.

Remark. Indications of this species from North Tian Shan (“ Kirghiski Ala tau, Ala Artscha, Frounze, 2500 m , juillet” – Verhulst, 1995 a: 303) are incorrect and I think that J.Verhulst seen a wrongly labelled or an aberrant of cocandica.

·       roschana Grum-Grshimailo, 1893  (Colias).  Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 27: 381. TL: “Koudara”. Had been originally described as a subspecies of alpherakyi. D: West Pamir .

·       usmatica Stshetkin, 1990  (Colias). Bulleten Moskovskogo Obstshestva ljubitelej prirody, Otdel biologicheskikh nauk, Moscow , 95: 59 – 60. TL: “западные отроги Туркестанского хребта: на склоне горного отрога Чукмартау близ посёлка Усмат, 2750 – 2800 м ” [western spurs of Turkestansky Mts.: on the slope of Chukmartau mountains near Usmat, 2750 – 2800 m]. Had been originally described as a subspecies of alpherakyi. D: Gissar.

 

aquilonaris Grum-Grshimailo, 1899  (Colias). Annales Musei Zoologici, Saint-Pétersbourg, 4: 457 - 458. TL: “F[luss] Olenek , 67 ½º B[reite]. 128 ¼º [sic] L[änge]” – by the lectotype (Grieshuber et al., 2001 : 170). Had been originally described as a variety of hecla.

·       aquilonaris Grum-Grshimailo, 1899 (Colias).  D: Chukotka; northern part of the Magadan region; north-west of East Siberia .

Remark. The specific level by J.Verhulst (1995 b).

 

chippewa Edwards, 1872  (Colias). Synopsis of the North American Butterflies, Washington : 9. TL: W and Great Slave L., Northwest Territories . Had been originally described as a good species.

·       gomojunovae Korshunov, 1996  (Colias). Dopolnenija i ispravlenija k knige “Dnevnye babochki asiatskoj chasti Rossii”, Novosibirsk : 18. TL: “Магаданская область, Билибинский район” [Magadan region, Bilibino district]. Had been originally described as a subspecies of palaeno. D: Chukotka; Magadan reg. (north).

 

cocandica Erschoff  in Fedtschenko, 1874 (Colias). Reise nach Turkestan, 2, 5 (3) (Lepidoptera): 6; pl. 1, fig. 3. TL: Stshurovsky gletsher, Turkestan Mts. Had been originally described as a variety of nastes.

·       cocandica Erschoff in Fedtschenko, 1874 (Colias). D: Tian Shan ; Darvaz; Gissar.

·       hinducucica Verity, [1911]  (Colias). Rhopalocera palaearctica, Firenze , [1]: 353; pl. 70, fig. 3. TL: “Ost Hinducuch”. Had been originally described as a race of cocandica. D: Pamir .

·       maja Grum-Grshimailo, 1891  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 25: 447. TL: “Thian Chan … Богусъ-Зуслунъ” [Bogus-Zuslun] – by the lectotype (Grieshuber & Churkin, 2003: 256). Had been originally described as a subspecies of cocandica. D: Dzhungaria.

·       mraceki Verhulst, 1999  (Colias). Lambillionea, Bruxelles, 99: 268 – 269, figs. 7, 8, 15, 16, 19. TL: “ Tadzhikistan , chaîne du Turkestan , col de Kumbel 2500 – 3000 m ”. Had been originally described as a subspecies of cocandica. D: Turkestan Mts.

·       nastoides Verity, [1911]  (Colias). Rhopalocera palaearctica, Firenze , [1]: 354. TL: “ Aram Koungei”, Transalai. Had been originally described in quadrinominal combination Colias cocandica cocandica nastoides. D: Alai and Transalai mountain ridges.

 

hecla Lefebvre, 1836  (Colias). Annales de la Société entomologique de France, Paris, 1836: 383; pl. 9B, fig. 3 – 6. TL: “Island” [ Greenland ]. Had been originally described as a good species.

·       orientis Wnukowsky, 1929  (Colias). Zoologische Anzeiger, 83: 221. TL: “In valle fl. Lenae et in valle fl. Olenek ”. Had been originally described as a variety of hecla. D: Polar tundras of Asiatic part and North Ural; Polar tundras of European part.

·       zamolodchikovi Churkin & Grieshuber, 2001 (Colias). Helios, Moscow , 2: 178 – 180; pl. 15, figs. 5, 6, pl. 16, figs. 1, 2. TL: “ Russia , East Chukotka, 20 km SE Ioni Lake , Gilmimlviem [65º 48’ N, 173º 22’ W]”. Had been originally described as a subspecies of hecla. D: Chukotka.

 

hyperborea Grum-Grshimailo, 1900  (Colias). Annales Musei Zoologici, Saint-Pétersbourg, 4: 455. TL: “Lena Thal, Wilui Mündung” – by the lectotype (Grieshuber et al., 2001: 169). Had been originally described as a good species.

·       hyperborea Grum-Grshimailo, 1900 (Colias). D: Chukotka; north of East Siberia .

·       paradoxa Churkin & Grieshuber, 2001 (Colias). Helios, Moscow , 2: 181 – 182; pl. 15, figs. 13 – 16. TL: “Sibiria, Putorana Plateau, Sibiria [sic!], Putorana Plateau [sic!], Eastern of Lama L., 200 m ”. Had been originally described as a subspecies of hyperborea. D: Putorana Plateau.

·       puella Churkin & Grieshuber, 2001 (Colias). Helios, Moscow , 2: 183 – 184; pl. 15, figs. 1 – 4. TL: “East Chukotka, Anadyr ’ vicinity”. Had been originally described as a subspecies of hyperborea. D: TL.

·       viluiensoides Verity, [1909]  (Colias). Rhopalocera palaearctica, Firenze , [1]: 250. TL: Srednekolymsk. Had been originally described as a race of hyperborea. D: Sayan; Praebaikal; Central Siberia .

 

jacutica Kurentzov, 1970  (Colias). Bulavousye cheshuekrylye Dalnego Vostoka SSSR, Leningrad : 35 – 36; ris. 28 (1), 29 (6). TL: “Якутия, р. Адыча” [Yakutia,  Adycha river (a tributary of Yana )] – by the lectotype (Churkin et al, 2001: 106 - 107). Had been originally described as a subspecies of nastes.

·       jacutica Kurentzov, 1970 (Colias). D: Yana and Indigirka valleys.

·       jacuticola Weiss & Mráček, 1989  (Colias). Nachrichten des entomologischen Vereins Apollo, N.F., Frankfurt am Main, 10: 282 – 287; fig. 1; pl. 1, 2, 3 [= yacuticola Verhulst, 1991 b: 36, syn.n. – unjustified emendation]. TL: “USSR, Siberia or., Yakutia mer. or., Oymyakon Valley, 50 km east of the village Tomtor, 500 – 600 m a.s.l.” Had been originally described as a good species. D: Suntar-Khajata Mts.

 

mongola Alphéraky, 1897  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 9: 188. TL: “Urga” – by the lectotype (Grieshuber & Churkin, 2003: 267). Had been originally described as a variety of nastes.

·       mongola Alphéraky, 1897 (Colias). D: South Transbaikal .

·       ukokana Korb & Yakovlev, 2000 (Colias).  Alexanor, Paris , 21 (1): 3 – 4; fig. 1. TL: “Altaï méridional, Oukok, rivére Ak-Alakha, 2 500 m ”. Had been originally described as a subspecies of Colias mongola. D: Altai (Kuraisky, Severo-Chujsky, Sajlugem and Katunsky mountain ridges; Ukok plateau); Tuva.

 

nastes Boisduval, 1832  (Colias). Icones historique des Lépidoptères nouveaux ou peu connus collections avec figures colorieers des papillons d’Europe nouvellement decouverts, Paris: pl. 8, fig. 4, 5. TL: [ Labrador ]. Had been originally described as a good species.

·       dezhnevi Korshunov  in Korshunov & Gorbunov, 1995 (Colias). Dnevnye babochki asiatckoj chastii Rossii, Ekaterinburg: 67 [nom. nov. pro Colias nastes sibirica Kurentzov, 1970, nec C. aurora sibirica Lederer, 1852]. TL: “Чапл[инские] кл[ючи], гор[ная] т[ундра]” [Chaplinskie Kluchi, Montainous tundra, East Chukotka ] – by the lectotype (Churkin et al, 2001: 105). Had been originally described as a subspecies of nastes. D: Northern part of East Siberia .

·       flinti Churkin, Grieshuber, Bogdanov & Zamolodchkov, 2001 (Colias). Helios, Moscow , 2: 108 – 109; pl. 10, figs. 11  - 15. TL: “ Russia , Wrangel I. , middle stream of Neizvestnaya R.”. Had been originally described as a subspecies of nastes. D: TL.

·       magadanica Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001 (Colias). Helios, Moscow , 2: 107 – 108; pl. 10, figs. 1 - 5. TL: “ Russia , Magadan region, Bol’shoi Annachag range, Aborigen v., 1200 m ”. Had been originally described as a subspecies of nastes. D: SE part of Magadan region.

·       nastes Boisduval, 1832 (Colias). D: North of the European part; North Ural; north of West Siberia .

·       streckeri Grum-Grshimailo, 1895  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 29: 290. TL: “in provincia Alberta ad Laggan collecta”. Had been originally described as a variety of nastes. D: Chukotka.

 

palaeno Linnaeus, 1761  (Papilio). Fauna Svecica, sistens animalia Sveciae Regni... (ed. altera): 272. TL: Sweden and Finland . Had been originally described as a good species.

·       arctica Nordström, 1927 (Colias). Arkiv for Zoologi, 19A (21): 3 – 4. TL: “Sredne-Kolymsk, Province Yakutsk, N.E.Siberia, Lat. 68º” – by the lectotype (Grieshuber & Worthy, 2004 b: 34). Had been originally described as a variety of palaeno. D: North of Euroasia

·       lapponica Staudinger, 1871  (Colias). Catalog der Lepidopteren des europaeischen Faunengebietes, Dresden (2 Aufl.): 6. TL: Lapland (northern part). Had been originally described as a variety of palaeno. D: Kola peninsula .

·       orientalis Staudinger, 1892  (Colias). Internationale entomologische Zeitschrift Iris , Berlin , 5: 310. TL: “Kentei, Amurgebiet”. Had been originally described as a variety of palaeno. D: all Siberia; all Far East without southern part of Ussuri and Amur districts.

·       palaeno Linnaeus, 1761  (Papilio).  D: Central European part.

 

tyche Boeber, 1812  (Papilio). Mémoires de la Société des Naturalistes de Mocsou, 3: 21; Tab. 1, fig. 3, 4. TL: “im südlichen Sibirien” (Grieshuber, 1998: 127). Had been originally described as a good species.

·       deckerti Verity, 1908  (Colias). Rhopalocera palaearctica, Firenze , [1]: pl. 42, fig. 27, 28. TL: “Monts Apfel et probablement aussi d’autres montagnes de cette région”. Had been originally described as a variety of melinos Eversmann, 1832. D: Transbaikal.

·       herzi Staudinger, 1901  (Colias). Catalog der Lepidopteren des palaearktischen Faunengebietes (ed. 1), Berlin : 16. TL: “Vilui”. Had been originally described as a variety of melinos Eversmann, 1847. D: Northern Siberia .

·       ludmilla Hemming, 1933  (Colias). Entomologist, London , 66: 278 [nom. nov. pro Colias tyche var. chryseis Verity, [1911], nec Colias erate f. chryseis Röber, 1907]. TL: “Amour”. Had been originally described as a race of melinos. D: Amur district.

·       montana Verity, [1911]  (Colias). Rhopalocera palaearctica, Firenze , [1]: 354. TL: “Montagnes de Tchuja, 1800 – 2400 m , S.-E. de l’Altai”. Had been originally described as a race of melinos. D: Altai; Tuva.

·       olga Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001 (Colias). Helios, Moscow , 2: 109 – 110;  pl. 9, figs. 1 – 5, 15. TL: “ Russia , East Chukotka, 20 km SE Ioni Lake , Gilmimlviem [65º 48’ N, 173º 22 W]”. Had been originally described as a subspecies of tyche. D: TL.

·       relicta Kurentzov, 1970  (Colias). Bulavousye cheshuekrylye Dalnego Vostoka SSSR, Leningrad : 38; ris. 29 (8). TL: “в горах истоков р. Охoты, где расположено несколько вершин  Верхоянского хребта, поднимающихся до 3000 м над уз.м.” [in mountains of sources of the Okhota river where are some summits of Verkhoyansky Mts. which have to 3000 m altitude]. Had been originally described as a subspecies of melinos. D: Far East ; Chukotka.

·       tyche Boeber, 1812  (Papilio). D: Sayan; Praebaikal.

·       werdandi Zetterstedt, 1840  (Colias). Insecta Lapponica: 908. TL: “In Lapponia septentrionali... in alpinum jugo latere Norvegico... Lapponia Scandinavica Torensis s. borealis”. Had been originally described as a subspecies of nastes. D: Northern part of European part and Ural.

 

Subgenus Neocolias Berger, 1986  

Lambillionea, Bruxelles, Suppl. 86: 7. TS: Papilio erate Esper, [1804].

 

aurorina Herrich-Schäffer, [1851]  (Colias). Systematische Bearbeitung der Schmetterlinge von Europa..., 6: 22. TL: Armenia . Had been originally described as a variety of thisoa.

·       aurorina Herrich-Schäffer, [1851] (Colias). D: Little Caucasus ; Transcaucasus.

·       daghestanica Verhulst, 1994  (Colias). Lambillionea, Bruxelles, 94: 115 – 117;  figs. 1, 2. TL: “Daghestan, Rutul 3000 m ”. Had been originally described as a subspecies of aurorina. D: Great Caucasus from Sulak river to Sumgaiti.

·       tamara Nordmann, 1851  (Colias). Bulletin de la Société impériale des Naturalistes de Moscou, 24: 413; Tab. 11, fig. 2, 3. TL: “Grusien”. Had been originally described as a good species. D: Great Caucasus from Maikop to Sulak river.

 

caucasica Staudinger, 1871  (Colias). Catalog der Lepidopteren des europaeischen Faunengebietes (ed. 2), Dresden : 6. TL: “Helenendorf” – by the lectotype (Nekrutenko, 1990: 99). Had been originally described as a subspecies of myrmidone.

Remark. S.Wagener (1990) listed the taxon balcanica Rebel (1901: 250) as a subspecies of caucasica.

·       anna Gerhard, 1882  (Colias). Berliner entomologische Zeitschrift, 26: 125. TL: “bei Achty am Samurflusse im Caucasus ”. Had been originally described as a variety of aurora. D: Great Caucasus (eastern part).

·       caucasuca Staudinger, 1871 (Colias). D: Little Caucasus from Alaverdi to Goradis; Armenia .

·       olga Romanoff, 1882  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 17: 127; pl. 4, fig. 1 – 4. TL: “aux environs de Borjom, d’Achalzich et d’Abastouman”. Had been originally described as a good species. D: Little Caucasus in Grusia.

 

chlorocoma Christoph, 1888  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 22: 308. TL: “Kasikoporan” – by the lectotype (Nekrutenko, 1990: 102). Had been originally described as a good species.

·       chlorocoma Christoph, 1888 (Colias). D: Nakhichevan region of Armenia .

·       tkatschukovi Bang-Haas, 1936  (Colias). Entomologische Zeitschrift, Frankfurt am Main , 50: 287. TL: “Russ. Armen. / Daralagaes / Pagod. Sultanbek” – by the lectotype (Nekrutenko, 1990: 102). Had been originally described as a subspecies of chlorocoma. D: NW Armenia .

 

erate Esper, [1801]  (Papilio). Die ausländischen oder die ausserhalb Europa zur Zeit in der übrigen Welttheilen vorgefundenen Schmetterlinge, Erlangen , 1: Tab. 119, Fig. 3. TL: South Russia [European part]. Had been originally described as a good species.

·       erate Esper, [1801] (Papilio). D: European part northwards to the Nizhny Novgorod region (1 male, 16.07.1988, N.Novgorod reg., Staraya Pustyn, without collector’s name; my collection); South Ural; plains of Kazakhstan, Usbekistan, Turkmenia and Kirghisia; Praealtai.

·       katboitzi Korb 2005 . (Colias) (fig. 6). Is described as a subspecies of erate.

Description. Male. Forewing length 25.0 – 26.5 mm . Upperside lemon yellow with a wide black marginal border; this border suffused with yellow scales. On the hindwing in the black marginal border have 1 – 5 yellow veins, on the forewing the yellow colouration of veins is rare. Forewing with black discal spot, hindwing with orange discal spot. Underside yellow, only in the anal part of the forewing the colouration is lighter, yellowish white. Underside with antemarginal row of dark points which are lighter on the forewing. Discal spot on the forewing with yellow centre, discal spot on the hindwing is silver. Female. Forewing length 23.0 – 26.0 mm . Ground colour as in male or white, with little basal dusting. Marginal border on the forewing with 2 – 6 yellow spots. On the hindwing this border fragmented. Underside as in male but in the white form the forewing colouration is whitish with 2 – 6 yellow spots. On the hindwing this border fragmented. Underside as in male but in the white form the forewing colouration is whitish with yellow apex.

Diagnose. This new subspecies distributes only in the Middle Asiatic part of Palaearctics. My materials was collected in the Kirghiz , Alai , Karzhantau, Transalai mountain ridges. I had be seen the specimens from Terskey Ala-Too, Transili Alatau, Talassky Ala-Too and Vanch Mts. too. I think that all others inhabite this subspecies. It differs from the nominotypical one by the less size (in ssp. erate forewing length is 19.0 – 21.0 mm in males). Females in both subspecies differ by the brighter colouration in the new one. Males of the new subspecies have bright black marginal border (in the nominotypical one this border is unbright, greyish black).

Type material. Holotype male, 26.06.1993, Kirghiz Mts., Chon-Kuurchak valley, 2600 m , S.Korb leg., deposite in the Zoological Institute, St.Petersburg. Paratypes: 10 males, 10 females, 26.06 – 10.07.1993, the same data as the holotype; 2 males, 2 females, 22 – 26.06.1992, Karzhantau Mts., A.Evdokimov leg.; 2 males, 2 females, 28.06.1997, Alai Mts., Kyzyl-Kyja, S.Korb leg.; 2 males, 3 females, 29.08.1998, Karakul Lake vicinity, G.Anufriev leg.; types are deposited in collections of Y.B.Kosarev, R.V.Yakovlev, A.G.Evdokimov, J.Grieshuber and Zoological Institute, St.Petersburg.

D: Middle Asiatic mountains.

 

myrmidone Esper, [1781]  (Papilio). Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen, Erlangen , 1: Tab. 65, Fig. 1, 2. TL: Hungary . Had been originally described as a good species.

·       antschari Korb 2005 . (Colias) (fig. 7). Is described as a subspecies of myrmidone. D: European part (without north and Ukraine – Moldova ).

·       ermak Grum-Grshimaïlo, 1890  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 301. TL: South Ural. Had been originally described as a variety of myrmidone. D: Middle and South Ural; southern parts of Tjumen and Kurgan regions.

·       myrmidone Esper, [1781] (Papilio). D: Moldova ; Ukraina.

 

polyographus Motschulsky, 1860  (Colias). Études d’Entomologie, 9: 29. TL: Japan: “ Hakodate ”. Had been originally described as a variety of neriene.

·       naucratis Fruhstorfer in Seitz, 1910  (Colias). Die Gross-Schmetterlinge der Erde, Stuttgart , 9: 165. TL: “Sibirien”. Had been originally described as a subspecies of hyale. D: South Praebaikal (Tajozhny); Krasnoyarsk vicinity.

·       polyographus  Motschulsky, 1860 (Colias). D: Transbaikal; southern Far East; Sakhalin ; Kuriles.

 

-?. sagartia Lederer, 1869  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 7: 75; pl. 4, fig. 1, 2. TL: Persia . Had been originally described as a good species.

Remark. This species is known from the following geographic points: “Iran du Nord, Elburs: Dizin Ouest de Gatchar 3000 – 3200 m , Kendevantunnel (N-O), 2900 – 3200 m , Mazandaran; Elburs Central: Demavend Ask 3600 m ; Sahand prés de Tabriz env. Lighwan 2500 – 3200 m ; Iran : Hadshyabad, Schahkuh” (Verhulst, 1993: 251). It’s a high montane inhabitant and can be found in Turkmenia, NW Kopet-Dagh.

 

Subgenus Eriocolias Berger, 1986  

Lambillionea, Bruxelles, Suppl. 86: 7. TS: Papilio chrysotheme Esper, [1781].

 

chrysotheme Esper, [1781]  (Papilio). Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen, Erlangen , 1: Tab. 65, Fig. 3, 4. TL: Hungary . Had been originally described as a good species.

·       audre Hemming, 1933  (Colias) (an bona species?). The Entomologist, London , 66: 278 [= andre Korshunov & Gorbunov, 1995: 67, syn.n.] [nom. nov. pro sibirica Grum-Grshimailo, 1893, nec sibirica Lederer, 1852]. TL: “Krasnojarsk”. Had been originally described as a subspecies of chrysotheme. D: Transural; South Siberia from Altai to Transbaikal.

·       chrysotheme Esper, [1781] (Papilio). D: European part in north to the Nizhny Novgorod region; Caucasus; Crimea; North Kazakhstan .

 

crocea Geoffroy  in Fourcroy, 1785 (Papilio). Entomologia Parisiensis: sive catalogus insectorum quae in agro Parisiensi reperiuntur; secundum methodum Geoffroeanum in sectiones, genera et species distributus; cui addita sunt nomina trivialia et fere trecentae  novae species, Parisiis: 250. TL: Paris . Had been originally described as a good species.

·       crocea Geoffroy in Fourcroy, 1785 (Colias). D: European part (without north); Caucasus; Transcaucasus; ?Altai; Crimea ; North and Middle Kazakhstan.

 

elena P.Gorbunov  in Korshunov & Gorbunov, 1995 (Colias). Dnevnye babochki asiatskoj chasti Rossii, Ekaterinburg: 67 – 68. TL: “Ю.-В. Алтай, п. Акташ” [SE Altai, Aktash]. Had been originally described as a subspecies of chrysotheme.

Remark. The specific rank by Korb, 1998a: 313.

·       elena P.Gorbunov in Korshunov & Gorbunov, 1995 (Colias). D: Central and South Altai .

 

eogene Felder & Felder, [1865]  (Colias). Reise der österreichischen Fregatte Novara, Lepidoptera: 196; Taf. 27, Fig. 7. TL: Himalayas . Had been originally described as a good species.

·       aphrodite Verity, 1907  (Colias). Rhopalocera palaearctica, Firenze , [1]: pl. 43, fig. 28 – 31. TL: “les montagnes du voisinage du fleuve Ak-sou ( Turkestan oriental, vers 44’ de lat et 80’ de long. E. de Greenwich”. Had been originally described as a variety of eogene. D: Gissar; Darvaz; Vanch Mts.

·       elissa Grum-Grshimaïlo, 1890  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 332. TL: “ Pamir septentr., Artscha Basch, Kara Kasouk”. Had been originally described as a forma of eogene. D: Alai and Transalai mountain ridges.

·       erythas Grum-Grshimaïlo, 1890  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 332. TL: “Hindou-Kouch or.”. Had been originally described as a forma of eogene. D: East Pamir .

·       theia Elwes, 1884  (Colias). Transactions of the Royal entomological Society of London, 32 (1): 13 - 14. TL: “…in the mountains near Osch, in Eastern Turkestan …”. Had been originally described as a good species. D: West Pamir .

 

erschoffi Alphéraky, 1881  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 16: 362; pl. 14, fig. 1, 2. TL: “Kouldjа”. Had been originally described as a good species.

·       erschoffi Alphéraky, 1881 (Colias). D: Dzhungaria; Ili mountains.

Remark. The locations of this species from “Tianschan Central”, “Turkestan, Alaп Mts, Ferghana, Burstan” and “ Koksu Range ” (Verhulst, 1991 a: 280) are incorrect.

 

fieldi Ménétriès, 1855  (Colias). Enumeratio corporum animalium musei imperialis Academiae Scientarum Petropolitanae, Petropoli, 1: 79; pl. 1, fig. 5. TL: “Monts Himalaya”. Had been originally described as a good species.

·       fieldi Ménétriès, 1855 (Colias). D: Southern Ussuri district (1 male, 6.09.1966, Zapovednik “Kedrovaja Pad”, A.V.Zvataev leg., coll. of the Zoological Museum of the Moscow University ).

Remark. The name chinensis Verity,1909  used in the book of Y.P.Korshunov & P.Y.Gorbunov (1995: 70) in the combination with Colias fieldi can’t be used for this species because this name published in the quadrinominal combination Gonepteryx rhamni nepalensis chinensis Verity, 1909: 284 (the subspecific status was given by Y.P.Nekrutenko (1968)) but not Colias.

 

heos Herbst  in Jablonsky, 1792 (Papilio). Natursystem der bekannten ausländischen Insekten, Schmetterlinge, 5: 213 – 214; Taf. 114, Fig. 5, 6. TL: “In Sibirien”. Had been originally described as a good species.

·       alpina Verity, [1911]  (Colias). Rhopalocera palaearctica, Firenze , [1]: 357; pl. 71, fig. 29, 30. TL: “Altai, Kentei et Changai”. Had been originally described as a forma of aurora. D: Altai; Tuva; Sayan; Praebaikal.

·       heos Herbst, 1792 (Papilio). D: Transbaikal; Amur district; Ussuri district.

 

ionovi  Korb, 2005. TL: Kirghiz Mts., Chon-Kuurchak valley, Say Lesnika, 2700 m . D: TL.

 

johanseni Troubridge & Philip, 1990  (Colias). Canadian Entomologist, 122: 16 – 18; figs. 1 – 3. TL: “ Canada , District of Mackenzie, Bernard Harbour , altitude 10 m ”. Had been originally described as a good species.

·       ssp? D: Chukotka.

 

marcopolo Grum-Grshimailo, 1888  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Peterburg, 32: 304. TL: “Hindukusch orient.” Had been originally described as a good species.

·       kushana Wyatt & Omoto, 1966  (Colias). Entomops, Nice, 5: 143 – 144; figs. TL: “Bala Quran, Anjuman Valley , Badachshan, 4,000 – 4,200 m ”. Had been originally described as a subspecies of marcopolo. D: South Pamir .

·       marcopolo Grum-Grshimailo, 1888 (Colias). D: East Pamir; West Pamir; Alai and Transalai mountain ridges.

 

regia Grum-Grshimailo, 1887  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 398. TL: “Transalai Fergana” – by the syntypes (Verhulast, 1997: 440). Had been originally described as a good species.

·       regia Grum-Grshimailo, 1887 (Colias). D: Alai and Transalai mountain ridges.

·       svetlanae Korb & Yakovlev, 1998  (Colias). Lambillionea, Bruxelles, 98: 140; fig. TL: “Tienschan, Inyltschek Tau Gebirge, Majdaadyr, 4000 m ”. Had been originally described as a subspecies of regia. D: All ridges of Inner Tian Shan.

 

romanovi Grum-Grshimailo, 1885  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 2: 229. TL: “Tekelik, Balakty, Katta-Karamuk”. Had been originally described as a good species.

·       romanovi Grum-Grshimailo, 1885 (Colias). D: North Tian Shan; Inner Tian Shan; Alai and Transalai mountain ridges.

 

sieversi Grum-Grshimailo, 1887  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 388, 397. TL: “Touptschek und Fuss der Kara Schar”. Had been originally described as a good species.

·       sieversi Grum-Grshimailo, 1887 (Colias). D: Zeravshansky, Gissarsky and Peter the Great mountain ridges (Verhulast, 1998: 214).

 

staudingeri Alphéraky, 1881  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Peterburg, 16: 368; pl. 14, fig. 3, 4. TL: “Tian-Chian, … Arschan” – by the lectotype (Grieshuber et al., 2004: 38 – 39). Had been originally described as a good species.

·       alexandrina Verity, 1908  (Colias). Rhopalocera palaearctica, Firenze , [1]: pl. 45, fig. 9. TL: “Les monts Alexandre”. Had been originally described as a variety of staudingeri. D: Kirghiz Mts; Kungey Ala Too Mts.; Transili Alatau Mts.; mountains southwards of Issyk Kul.

·       maureri Staudinger, 1901  (Colias). Catalog der Lepidopteren des palaearktischen Faunengebietes (ed. 3), Drezden: 18. TL: “Ferg. m. (Transalai)”. Had been originally described as a variety of staudingeri. D: Alai Mts. (western part).

·       pamira Grum-Grshimaïlo, 1890  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 334; pl. 4, fig. 2. TL: “les confins des elevations du Pamir , seulement sur les versant N.O. de la caine transalaienne”. Had been originally described as a variety of staudingeri. D: Transalai Mts.; Alai Mts. (western part); West Tian Shan; Inner Tian Shan up to line western shore of Issyk Kul – Kashi.

·       staudingeri Alphéraky, 1881 (Colias). D: Terskey Ala Too Mts. in the Karakol vicinity.

 

sulitelma Aurivillius, 1890  (Colias). Bih. K. Svenska Vetensk-Akademiae Handliae, 15 (4) (1): 12; pl. 2, fig. 10. TL: Sweden , Sulitelma Mt. Had been originally described as a subspecies of hecla.

·       kurnakovi Kurentzov, 1970  (Colias). Bulavousye cheshuekrylye Dalnego Vostoka SSSR, LeningraD : 40 – 41; ris. 32 (1). TL: “в горных тундрах Омсукчанского хребта Магаданской обл.” [in the montane tundra of Omsukchansky Mts. in the Magadan region]. Had been originally described as a subspecies of  sulitelma. D: North Far East ; Chukotka.

·       sulitelma Aurivillius, 1890 (Colias). D: Tundras of European part and Siberia .

 

thisoa Ménétriès, 1832  (Colias). Catalogue raisonné des objets de zoologie recueillis dans un voyage au Caucase et jusqu’alors frontiиres actuelles de la Perse , enterpris par ordre de S.M.Empereur, Saint-Pétersbourg: 244. TL: Shakhdagh Mt. , Caucasus – by the lectotype designation (Nekrutenko, 1990: 98). Had been originally described as a good species.

·       aeolides Grum-Grshimaïlo, 1890  (Colias). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 346 - 347. TL: “montagnes alaïennes, dans le Trans-Alaï et les monts de Pierre le Grand”. Had been originally described as a variety of thisoa. D: Inner Tian Shan; Alai ; Darvaz.

·       irtyschensis Lukhtanov, 1999 (Colias).  Atalanta, Würzburg, 30 (1/4): 137 – 138. TL: “Kasachstan, Ust-Kamenogorsk-Gebiet, Altai, Kalba-Gebirge, Tainty.” Had been originally described as a subspecies of thisoa. D: SW Altai: mountain ridges Kurchumsky, Kalbinsky, valley of the Irtysh river near Bukhtarma; possible that this subspecies flies in Saur and Tarbagatai too.

·       nikolaevi Korshunov, 1998 (Colias).  Novye opisanija i utochnenija dlja knigi “Dnevnye babochki asiatskoj chasti Rossii”, Novosibirsk : 14. TL: "Алтай, Курай, урочище Тюргуно, степной участок" [Altai, Kurai, Tyurguno valley, steppe area]. Had been originally described as a subspecies of thisoa. D: Altai (without SW); Sayan; Tuva; Transbaikal.

·       strandiana Sheljuzhko, 1935  (Colias). Folia zoologia hydrobiologia, Riga , 8: 138. TL: “ms. Aragлs / (Gub. Erivanj)” – by the lectotype (Nekrutenko, 1990: 98). Had been originally described as a subspecies of thisoa. D: Small Caucasus ; Transcaucasus; Kopet-Dagh.

·       thisoa Ménétriès, 1832 (Colias). D: Great Caucasus .

·       urumtsiensis Verity, 1909  (Colias). Rhopalocera palaearctica, Firenze , [1]: 252. TL: Urumchi. Had been originally described as a race of thisoa. D: West and North Tian Shan ; Dzhungaria.

 

viluiensis Ménétriès, 1859  (Colias). Bulletin de l’Academie Scientifique, Saint-Pétersbourg, 17: 213. TL: “Vilui”. Had been originally described as a good species.

·       heliophora Churkin & Grieshuber, 2001 (Colias). Helios, Moscow , 2: 175 – 177; pl. 15, figs. 11, 12, pl. 16, figs. 4, 5. TL: “Chukotka, Bilibino, B. Koperveem R., 350 m ”. Had been originally described as a subspecies of viluiensis. D: Chukotka.

·       viluiensis Ménétriès, 1859 (Colias). D: Putorana plateau; Yakutia; Magadan reg.; Middle Far East.

 

wiskotti Staudinger  in Staudinger & Bang-Haas, 1882 (Colias). Berliner entomologische Zeitschrift, 26: 166; Taf. 2, fig. 9, 10. TL: “ Samarkand ”. Had been originally described as a good species.

·       aurea Kotzsch, 1936  (Colias). Entomologische Rundschau, 54 (5): 45. TL: “Ost-Hindukusch, Alpenwiesenzone, 3500 bis 3800 m ”. Had been originally described as a subspecies of wiskotti. D: West Pamir ; Darvaz.

·       chrysoptera Grum-Grshimailo, 1888  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 22: 305. TL: “Roschan ( Afghanistan ), Transalai (var?), Sarykol”. Had been originally described as a variety of wiskotti. D: East Pamir; Transalai Mts. (eastern part).

·       draconis Grum-Grshimailo, 1891  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 25: 465. TL: “Kisyl-Art Mt. Alexandr” (by the lectotype – Grieshuber & Worthy, 2004 a: 28); “Ошибочно показана мною водящейся на Памире, в действительности она была найдена в горном районе к западу от оз. Иссык-Куля” [Erroneously stated by me to occur in Pamir, in fact it was found in a montaneous region westwards of Issyk Kul] (Grum-Grshimailo, 1907: 430, remark; Tshikolovets & Kostjuk, 1994: 16 – english translation). D: West Tian Shan : Talassky and Chatkalsky mountain ridges.

·       rueckbeili O.Bang-Haas, 1927  (Colias). Horae Macrolepidopterologicae regionis Palaearcticae, Drezden, 1: 113 [nom. nov. pro alexandrina Röber, 1907, nec alexandrina Edwards, 1863]. Had been originally described as a variety of wiskotti. D: Kirghiz Mts. (west).

·       separata Grum-Grshimailo, 1888  (Colias). Horae Societatis Entomologicae Rossicae, Sankt-Pétersburg, 22: 305. TL: “Montes Alaienses”. Had been originally described as a variety of wiskotti. D: Alai and Transalai mountain ridges.

·       seres Groum-Grshimaïlo, 1890 (Colias).  Mémoires sur les Lépidoptères, Saint-Pétersbourg,  4: 352 – 353. TL: “de Goudjabaï, sur les pentes N. E. des monts Kounjout”. Had been originally described as a variety of wiskotti. D: Eastern parts of Muzkol and Vakhansky Mts.

·       wiskotti Staudinger  in Staudinger & Bang-Haas, 1882 (Colias). D: Gissar.

 

Tribus Callidryini  Kirby, 1896

 

Zoology in illustrations, London , 1: pl. 22. TS: Papilio hecabe Linnaeus, 1758.

 

hecabe Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 470. TL: China . Had been originally described as a good species.

·       ssp? 1 male, 08.1969, South Ussuri region, remaining data unknown; this specimen was collected by Dr. G.A.Anufriev during his work about the Far Eastern fauna of cicads and deposite in the collections of the Zoological Museum of Nizhny Novgorod university.

 

Brewster’s Edinburgh . Encyclopedia, 9: 127. TS: Papilio rhamni Linnaeus, 1758.

 

aspasia Ménétriès  in Schrenck, 1859 (Gonopteryx). Reisen und Forschungen im Amurlande in den Jahren 1859 – 1867, St .-Petersburg, 2: 17; Taf. 1, Fig. 8. TL: “Amur”. Had been originally described as a good species.

·       aspasia Ménétriès in Schrenck, 1859 (Rhodocera). D: Amur district; Ussuri district.

 

farinosa Zeller, 1847  (Gonepteryx). Isis, von Ocken, 1: 5. TL: Fetchie, Mugla , Turkey . Had been originally described as a good species.

·       farinosa Zeller, 1847 (Gonepteryx). D: Transcaucasus; Kopet Dagh; Gissar; Darvaz; Alai; West Pamir .

 

maxima Butler , 1885 (Gonepteryx). Annales of the Magazine of natural History, London , 5 (15): 407 - 408.  TL: Japan , “ Nikko ”. Had been originally described as a good species.

·       amurensis Graeser, 1888 (Gonepteryx) . Deutsche entomologische Zeitschrift Iris, 32: 70. TL: “Chab.[arowsk]”. Had been originally described as a variety of rhamni. D: Amur district (from Blagovestshensk to Troizkoe); Ussuri district.

Remark. Y.P.Korshunov (1998: 13) wrote: “В книге московских коллекционеров "Butterflies of Russia", v. 1, 1997 г . для окр. Хасана указан Gonepteryx amintha limonia Mell, 1943. Экземпляр на таблице 40 этой книги под номером 15 скорее всего (если этикетка верна) аберрация maxima, от которой он отличается лишь некоторым налетом красноватых чешуек на п. кр. Какова же amintha? Это один из самых крупных в роде видов (размах крыльев 65 – 80 мм ). Что касается подвида limonia, то это вообще "Größte Form" , как стоит в описании, и с далеким южно-китайским ареалом. Ближе к номеру 15 бабочки подвида thibetana Nekrutenko, 1968 , но это из Тибета, что очень далеко от Хасана" [In the book of the Moscow collectors "Butterflies of Russia", v. 1, 1997 for the Khasan vicinity is specified Gonepteryx amintha limonia Mell, 1943 . A specimen on the table 40 of this book under number 15 most likely (if the label is correct) an aberratio of  maxima, from which it differs only by some tinge of reddish scales on the forewing. What amintha? It is one of largest in a genus of species (scope of wings 65 – 80 mm ). As to подвида limonia, it in general "Gröβste Form", as stands in the description, and with far South-Chinese area. Closer to number 15 butterflies of subspecies thibetana Nekrutenko, 1968, but it from Tibet , that very much far from Khasan].

 

rhamni Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 470. TL: Sweden . Had been originally described as a good species.

·       miljanowskii Nekrutenko, 1966  (Gonepteryx). Zeitschrift der Wiener entomologischen Gesellschaft, 51: 45; Taf. 3, Fig. 1 – 3, 6 – 8. TL: “Suchumi”. Had been originally described as a subspecies of rhamni. D: Small Caucasus ; Transcaucasus.

·       nana Wnukowsky, 1935  (Gonepteryx). Lambillionea, Bruxelles, 6: 131. TL: Altai. Had been originally described as a variety of rhamni. D: Southern part of West Siberia ; Altai.

·       rhamni Linnaeus, 1758 (Papilio). D: North and central European part; Carpathians.

·       tianschanica Nekrutenko, 1970  (Gonepteryx). The Journal of the Lepidopterist’s Society, 24: figs. 1 – 6. TL: “ Union of Soviet Socialist Republics , Pravy Talgar, Tian Shan Mts., 1500 m , Alma-Ata prope” Had been originally described as a subspecies of rhamni. D: North Tian Shan ; Dzhungaria.

·       transiens Verity, 1913  (Gonepteryx). Journal of the Linnean Society (Zoology), 32: 180. TL: Italy . Had been originally described as a subspecies of rhamni. D: Southern European part; Great Caucasus; Crimea; North Kazakhstan .

 

Subfamily Anthocharinae  Scudder, 1889

Tribus Anthocharini  Scudder, 1889

 

Coll. icon. hist. Chenil. Europ., 21: pl. 5. TS: Papilio cardamines Linnaeus, 1758.

 

cardamines Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 468. TL: Sweden . Had been originally described as a good species.

·       alexandra Hemming, 1933 (Anthocharis). Entomologist, London , 66: 277. TL: Kirghiz Mts. Had been originally described as a subspecies of cardamines.  D: All mountains of Middle Asia without Pamir .

·       callisto Churkin, 2004 (Anthocharis). Helios, Moscow , 5: 133 – 135; pl. 10, figs. 5 – 8. TL: “Tadjikistan, Peter the Great Range , 23 km SE Tadzhikobad [sic!], Ganishou v.”. Had been originally described as a subspecies of cardamines. D: TL.

·       cardamines Linnaeus, 1758 (Papilio). D: European part; Caucasus; Transcaucasus; Crimea; North Kazakhstan .

·       bambusaroides Huang & Murayama, 1992  (Anthocaris). Tyô to Ga, Tokyo , 43: 4; figs. 9, 10. TL: “Halashi, Altai, 1300 – 1700 m , grassland in forest zone”. Had been originally described as a subspecies of cardamines. D: SW Altai.

·       kobayashii Matsumura, 1925 (Anthocharis). Journal of the College of Agriculture of Hokkaido Imperial University in Sapporo , 15 (3): 91.  TL: North Sakhalin : “Onory”.  Had been originally described as a subspecies of cardamines. D: Sakhalin .

·       progressa Sovunsky, 1905  (Euchloe). Revue Russe d’Entomologie, Moscow , 5: 103. TL: Irkutzk. Had been originally described as a variety of cardamines. D: Siberia; Far East .

 

damone Boisduval, 1836  (Anthocharis). Histoire naturelle des insectes. Species général des Lépidoptéres, Paris, 1: 564. TL: Sicily . Had been originally described as a good species.

·       eunomia Freyer, [1851]  (Anthocharis). Neue Beiträge zur Schmetterlingskunde mit Abbildungen nach der Natur, Augsburg , 6: 149; Taf. 547, fig. 3. TL: Turkey . Had been originally described as a subspecies of damone. D: South Transcaucasus .

 

gruneri Herrich-Schäffer, 1845  (Euchloe). Systematische Bearbeitung der Schmetterlinge von Europa..., Regensburg , 1: Taf. 551, Fig. 4. TL: Amasia in Turkey (Reissinger, 1989: 62). Had been originally described as a good species.

·       armeniaca Christoph, 1893  (Anthocharis). Deutsche entomologische Zeitschrift Iris, 6: 86. TL: “Ordub.[ad]” – by the lectotype (Nekrutenko, 1990: 71). Had been originally described as a subspecies of gruneri. D: Transcaucasus.

 

Nasekomye cheshuekrylye (Insecta, Lepidoptera), Leningrad , 1 (2): 126. TS: Anthocharis scolymus Butler , [1866]

 

scolymus Butler , [1866]  (Anthocharis). Proceedings of the Linnean Society of London , Zoology, 9: 52. TL: Japan , Hokkaido , Hakodade. Had been originally described as a good species.

·       mandschurica O.Bang-Haas, [1930] (Midea).  Die Gross-Schmetterlinge der Erde, Supplement 1: 105. TL: Charbin in N China . Had been originally described as a subspecies of scolymus. D: Ussuri district northwards to Khabarovsk .

 

Verzeichniss bekannter Schmett[er]linge, Augsburg , 6: 94. TS: Euchloe ausonia Hübner var. esperi Kirby, 1871.

 

ausonia Hübner, [1803]  (Papilio). Sammlung europäischer Schmetterlinge, Augsburg , 1: Fig. 582, 583. TL: North Italy . Had been originally described as a good species.

·       dubatolovi Korshunov  in Korshunov & P.Gorbunov, 1995 (Euchloe). Dnevnye babochki asiatskoj chasti Rossii, Ekatheringurg: 62. TL: “Алтай, Катунский хр., 15 км ЮЗ с. Катанды, между реками Кураган и Кучерла, альпийский луг, 2300 м ” [Altai, Katunsky Mts., 15 km SW of Katandy, between rivers Kuragan and Kucherla, alpine meadow, 2300 m]. Had been originally described as a subspecies of ausonia. D: Altai.

·       pulverata Christoph, 1884  (Anthocharis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 1: 99. TL: “Askhabad, Kysyl-Arvat”. Had been originally described as a variety of belia Linné, 1767. D: Deserts of Middle Asia and South Kasakhstan .

·       transiens Verity, 1908  (Euchloe). Rhopalocera palaearctica, Firenze , [1]: 180. TL: Alai valley. Had been originally established in the quadrinominal combination Euchloe belia simplonia race transiens. D: Mountains of Tian Shan, Alai , Gissar and Darvaz.

·       volgensis Krulikowsky, 1897  (Euchloe). Trudy Saratovskogo obstshestva estestvoispytatelej i ljubitelej prirody, Saratov , 1: 36. TL: Saratov . Had been originally described as a variety of belia Linné, 1767. D: South of European part; Caucasus; Transcaucasus; Kopet-Dagh; South Ural; Crimea .

 

creusa Doubleday & Hewitson, [1847]  (Anthocharis). Genera of the diurnal Lepidoptera, 1: pl. 7, fig. 1. TL: “Rocky Mountains”, N. America . Had been originally described as a good species.

·       emiorientalis Verity, [1911]  (Euchloe). Rhopalocera palaearctica, Firenze , [1]: 338. TL: “Altai, Ongodai”. Had been originally established in the quadrinominal combination Euchloe belia creusa emiorientalis. D: Altai; Tuva; Sayan; Praebaikal.

·       kurentzovi Beljaev, 1986  (Euchloe). Sistematika i ekologia cheshuekrylykh Dalnego Vostoka SSSR, Vladivostok : 117; ris. 6, 9, 10. TL: “Магаданская  обл., Омсукчан” [Magadan reg., Omsukchan]. Had been originally described as a subspecies of orientalis. D: North Siberia; Chukotka; Far East .

·       nemoralis Beljaev, 1986  (Euchloe). Sistematika i ekologia cheshuekrylykh Dalnego Vostoka SSSR, Vladivostok : 117 – 118; ris. 11, 12. TL: “Приморский край, Спасский р-н, Кнорринг” [Ussuri district, Spassk area, Knorring]. Had been originally described as asubspecies of orientalis. D: Ussuri district.

·       orientalis Bremer, 1864  (Anthocharis). Mémoires de la Academie impériale des Sciences, Saint-Pétersbourg, 8: 8. TL: Transbaikalia: “Onon”. Had been originally described as a variety of belemnida Hübner, [1819]. D: Transbaikal; Amur district.

 

charlonia Donzel, 1842  (Anthocharis). Annales de la Société entomologique française, Paris, 11: 197; pl. 8, fig. 1. TL: “Emsilah en Barbarie”. Had been originally described as a good species.

·       charlonia Donzel, 1842 (Anthocharis). D: Kopet Dagh; deserts of Gissar.

 

daphalis Moore , 1865  (Euchloe). Proceedinds of the Zoological Society of London , 1865: 491, pl. 31, fig. 14.  TL: “Middle Kunawur”. Had been originally described as a good species.

·          daphalis Moore , 1865 (Euchloe). D: West Pamir .

 

simplonia Freyer, 1829  (Pontia). Beiträge zur Geschichte europäischer Schmetterlinge mit Abbildungen nach der Natur, Augsburg , 2: pl. 73, fig. 2. TL: Alps . Had been originally described as a good species.

·       jakutia Back, 1990  (Euchloe). Atalanta, Würzburg, 21: 193 – 195; Taf. 2, Abbn. 3, 4. TL: “ USSR , Yakutia, Suntar Mts. ” Had been originally described as a subspecies of simplonia. D: Far East ; Chukotka.

·       kusnezovi Korshunov  in Korshunov & P.Gorbunov, 1995 (Euchloe). Dnevnye babochki asiatskoj chasti Rossii, Ekatherinburg: 63. TL: “низовья Енисея, среднее течение р. Ирбо-Кета” [under course of Enisej, middle course of Irbo-Keta]. Had been originally described as a subspecies of naina. D: North Siberia .

Remark. In the synonymic list of V.K.Tuzov (1993: 18) from “N.Siberia” listed the taxon occidentalis Verity, 1908, but this subspecies don’t can to inhabite the North Siberian territory because “...Verity (1908: XXXI) listed distribution: southern and western Mediterranean, as rece also in Syria and race occidentalis has distribution N.Italy, France, Spain” (Kudrna, 1983: 59); as well as this taxon is a synonym of Euchloe crameri crameri Butler, 1869 (partim), E. c. esperi Kirby, 1871 (partim) and E. c. melanochloros Röber, 1907 (partim) (Reissinger, 1989: 157).

·       naina Kozhantschikov, 1923  (Euchloe). Jahrbuch des Martjanower Staatsmuseum, Minussinsk, 1: 3. TL: “am Buja See im Sajan-Gebirge”. Had been originally described as a subspecies of belia. D: Altai; Sayan; Tuva; Praebaikal; Transbaikal; Amur district.

 

 

Histoire naturelle des insectes. Species général des Lépidoptères, Paris, 1: 552. TS: Papilio eupheme Esper, [1805].

 

eupheme Esper, [1805]  (Papilio). Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen, Erlangen , Suppl. 1 (1): 105; Tab. 113, Fig. 5. TL: Sevastopol . Had been originally described as a good species.

·       eupheme Esper, [1805] (Papilio). D: South of European part; northern slope of Great Caucasus; deserts of Kazakhstan and Turkestan; Tian Shan; Dzhungaria; Crimea; Altai: Barnaul vicinity (Yakovlev, 2002: 174).

·       menestho Ménétriès, 1832  (Pieris). Catalogue raisonné des objets de zoologie recueillis dans un voyage au Caucase et jusqu’alors frontiиres actuelles de la Perse , enterpris par ordre de S.M.Empereur, Saint-Pétersbourg: 245. TL: “Mont. Taliisch” – by the lectotype (Nekrutenko, 1990: 75). Had been originally described as a good species. D: Southern slope of Great Caucasus; region between Kura and Araks rivers; Little Caucasus; Transcaucasus.

 

fausti Christoph, 1877  (Zegris). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 12: 231; pl. 5, fig. 1, 2. TL: Krasnovodsk. Had been originally described as a good species.

·       fausti Christoph, 1877 (Zegris). D: Deserts of Middle Asia .

 

Études de Lépidoptèrologie, 7: 232. TS: Pontia pyrothoe Eversmann, 1832.

 

pyrothoe Eversmann, 1832  (Pontia). Mémoires de la Société des Naturalistes de Moscou, 8: 352; pl. 20, fig. 3, 4. TL: “in monticulis Inderiensibus ad Jaicum interiorem” [Indersk vicinity]. Had been originally described as a good species.

·       pyrothoe Eversmann, 1832 (Pontia). D: Southern Volga region; South Praeural; Kazakhstan ; deserts of Middle Asia.

 

Subfamily Pierinae  Duponchel, [1835]

Tribus Teracolini  Reuter, 1896

 

Verzeichniss bekannter Schmett[er]linge, Augsburg , 7: 97. TS: Papilio amata Fabricius, 1775.

 

fausta Olivier, [1804]  (Papilio). Voyage dans l’Empire Othoman, 4: 29; pl. 33, fig. 4 A , B. TL: Beirout. Had been originally described as a good species.

·       fausta Olivier, [1804] (Papilio). D: Kopet Dagh; South Turkmenia; South-West Tadzhikistan .

 

Tribus Pierini  Duponchel, [1835]

 

Verzeichniss bekannter Schmett[er]linge, Augsburg , 6: 90. TS: Papilio crataegi Linnaeus, 1758.

 

crataegi Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 467. TL: Sweden . Had been originally described as a good species.

·       adherbal Fruhstorfer, 1910 (Aporia) . Societas Entomologica, 25: 50. TL: Japan: “Yesso, Sapporo ”. Had been originally described as a subspecies of crataegi. D: Kuriles.

·       crataegi Linnaeus, 1758 (Papilio). D: European part; Crimea; Kazakhstan (without mountains); Caucasus ; Transcaucasus; Kopet Dagh.

·       meinhardi Krulikowsky, 1909  (Aporia). Revue Russe d’Entomologie, Moscow , 9: 270. TL: Tomsk . Had been originally described as an aberratio of crataegi. D: All Asiatic part of Russia .

·       sachalinensis Matsumura, 1925  (Aporia). Journal of the College of Agriculture of Hokkaido Imperial University of Sapporo , 15 (3): 90, pl. 10, fig. 10. TL: Sakhalin : “Odomari, Ichinosawa, Kaizuka, Toyohara, Konuma, Pilwo, Onory, Alexandrowsk”. Had been originally described as a subspecies of crataegi. D: Sakhalin .

·       shugnana Sheljuzhko, 1925  (Aporia). Mitteilungen der Münchener entomologische Gesellschaft, 15: 97. TL: Shugnan Mts. Had been originally described as a subspecies of crataegi. D: West Pamir .

·       tianschanica Rühl, [1893]  (Aporia). Die palaearktischen Grossschmetterlinge und ihre Naturgeschichte. Tagfalter, Leipzig , 1: 231. TL: Tian Shan . Had been originally described as a variety of crataegi. D: All Middle Asiatic mountains without Pamir .

 

hippia Bremer, 1861 (Pieris) . Bulletin de la Academie des Sciences, Saint-Pétersbourg, 3: 543. TL: “von der Dzeja bis zur Bureja-Gebirge”. Had been originally described as a good species.

·       hippia Bremer, 1861 (Pieris). D: Amur district; Ussuri district.

·       occidentalis O.Bang-Haas, 1927  (Aporia). Horae Macrolepidopterologicae regionis palaearcticae, Dresden , 1: 107. TL: “Kiachta”. Had been originally described as a subspecies of hippia. D: Transbaikal.

 

Cistula Entomologica, 1: 38, 51. TS: Pontia leucodice Eversmann, 1843.

 

illegiblata Korb, 1998  (Metaporia). Lambillionea, Bruxelles, 98: 469 – 470; Abbn. 3, 4, 6. TL: “Kirgisien, Terskey Ala-Too-Gebirge, Kadshi-Saj”. Had been originally described as a good species.

·       illegiblata Korb, 1998 (Metaporia). D: Kirghiz Mts. ; Terskey Ala Too Mts.; Saush-Khozor valley in Tadzhikistan .

 

leucodice Eversmann, 1843  (Pontia). Bulletin de la Société des Naturalistes de Moscou, 3: 541; pl. 7, fig. 2. TL: “Noor-Saisan”. Had been originally described as a good species.

·       aryana Wyatt & Omoto, 1966  (Metaporia). Entomops, Nice, 5: 149 – 150; figs. TL: “Parian, Panjshir Valley , Hindu-Kush, 2,500 m ”. Had been originally described as a subspecies of leucodice. D: Darvaz; Pamir .

·       illumina Grum-Grshimaïlo, 1890  (Metaporia). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 155. TL: Peter the Great Mts. Had been originally described as a variety of leucodice. D: All Middle Asiatic mountains without Darvaz and Pamir .

·       leucodice Eversmann, 1843 (Pontia). D: Dzhungaria; Tarbagatai.

·       morosevitshae Sheljuzhko, 1907  (Metaporia). Revue Russe d’Entomologie, Moscow , 7: 233. TL: “Karatau”. Had been originally described as a subspecies of leucodice. D: West Tian Shan .

 

Verzeichniss bekannter Schmett[er]linge, Augsburg , 3: 42. TS: Papilio mesentina Cramer, 1777.

 

mesentina Cramer, 1777  (Papilio). Uitleven Kapellen, 2 (14): 102; tab. 162, fig. B, C. Had been originally described as a good species.

·       turanica Sheljuzhko, 1908  (Pontia). Revue Russe d’Entomologie, Moscow , 7: 233. TL: Kopet Dagh. Had been originally described as a variety of mesentina. D: South Turkmenia; Kopet Dagh; South Gissar .

Fauna Boica, 2 (1): 152, 161. TS: Papilio brassicae Linnaeus, 1758.

 

brassicae Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 467. TL: Sweden . Had been originally described as a good species.

·       brassicae Linnaeus, 1758 (Papilio). D: European part; Caucasus; Transcaucasus; Kazakhstan; Crimea; Siberia (southern and middle part of Baikal); Amur and Ussuri districts (Korshunov & Gorbunov, 1995; Streltsov, 1997); Kopet Dagh.

·       ottonis Röber  in Seitz, 1907 (Pieris) ( [3] ). Die Groβschmetterlinge der Erde. Die Groβschmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart , 1: 45. TL: “ Fergana ”. Had been originally described as a subspecies of brassicae. D: All mountains of Middle Asia.

 

deota de Nicéville, [1884]  (Mancipium). Journal of the Asiatic Society, Bengal , 7 (2): 82; pl. 9, fig. 10. TL: “at Gya, Ladak”. Had been originally described as a good species.

·       deota de Nicéville, 1884 (Mancipium). D: Pamir; Transalai Mts. (Lukhtanov, 1996: 212).

 

eitschbergeri Lukhtanov, 1996  (Pieris). Atalanta, Würzburg, 27: 213 – 218;  Textabbn. 1, 3 – 5, 9, 11; Taf. 5, Abbn. 1, 2. TL: “Kirgisien, Inner-Tienschan, Stromgebiet des Flusses Naryn, Dshetim-Gebirge, 70 km E. Naryn, 2900 – 3400 m ”. Had been originally described as a good species.

·       eitschbergeri Lukhtanov, 1996 (Pieris). D: Dzhetim Mts. ; Naryntau Mts. ; Issyk Kul region; Kirghiz Mts.

 

tadjika Grum-Grshimailo, 1888  (Pieris). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 22: 304. TL: “Darwaz-Kette, Kalai-Chum” (Grum-Grshimailo, 1887: 379); “Darwaz, Karategin” (Grum-Grshimailo, 1888: 304). Had been originally described as a good species.

·       tadjika Grum-Grshimailo, 1888 (Pieris). D: Darvaz; West Pamir

Les variations geographiques et saisonnieres des papillons diurnes en France, Paris, 1: 121. TS: Papilio napi Linnaeus, 1758 (Kudrna, 1974; Kudrna & Geiger, 1985: 55).

 

banghaasi Sheljuzhko, 1910  (Pieris). Revue Russe d’Entomologie, Saint-Pétersbourg, 9: 385. TL: Naryn. Had been originally described as a variety of napi.

·       banghaasi Sheljuzhko, 1910 (Pieris). D: Dzhungaria; Inner Tian Shan .

·       kirgizskana Korb, 2001 (Artogeia). Alexanor, Paris , 21 (3): 151 – 152. TL: “Kirghizie, chaîne Kirghize, vallée du Tchon-Kouourtchak, 2 500 m ”. Had been originally described as a subspecies of banghaasi. D: North Tian Shan .

 

bowdeni Eitschberger, [1984]  (Pieris). Herbipoliana, Marktleuthen, 1: 218. TL: Amasia in Turkey . Had been originally described as a good species.

·          bowdeni Eitschberger, [1984] (Pieris). D: Southern Transcaucasus .

 

bryoniae Hübner, [1806]  (Papilio). Sammlung europäischer Schmetterlinge, Augsburg , 1: Fig. 62. TL: “aus den Gebirgen bei Gentf”. Had been originally described as a good species.

·       bryonides Sheljuzhko, 1910  (Pieris). Revue Russe d’Entomologie, Saint-Pétersbourg, 9: 384. TL: “Lepsinsk”. Had been originally described as a variety of napi. D: Dzhungaria; North Tian Shan ; Tarbagatai; Altai; Sayan.

·       caucasuca Verity, 1908  (Pieris). Rhopalocera palaearctica, Firenze , [1]: 144. TL: region of Kuban . Had been originally established in the quadrinominal combination Pieris napi bryoniae caucasica. D: Caucasus ; Transcaucasus.

·       kamtschadalus Röber  in Seitz, 1907 (Pieris). Die Groβschmetterlinge der Erde. Die Groβschmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart , 1: 49. Had been originally described as a variety of napi. TL: “Kamtschatka”. D: Kamchatka .

·       lappona Rangnow, 1935  (Pieris). Entomologische Rundschau, 52: 175 – 176. TL: “Lule, Lappmark” – by the lectotype (Eitschberger, 1989: 207). Had been originally described as a subspecies of napi. D: North of European part.

·       schintimeisteri Eitschberger, [1984]  (Pieris). Herbipoliana, Marktleuthen, 1: 129. TL: Aldan river: Tommot. Had been originally described as a subspecies of bryoniae. D: Far East .

·       sheljuzhkoi Eitschberger, [1984]  (Pieris). Herbipoliana, Marktleuthen, 1: 128. TL: Magadan region, Omsukchan. Had been originally described as a subspecies of bryoniae. D: Chukotka; northern part of Magadan region.

·       sifanica Grum-Grshimailo, 1895  (Pieris). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 29: 290. TL: Amur. Had been originally described as a subspecies of napi. D: Amur district.

·       vitimensis Verity, [1911]  (Pieris). Rhopalocera palaearctica, Firenze , [1]: 332. TL: Vitim . Had been originally established in the quadrinominal combination Pieris napi frigida vitimensis. D: Transbaikal.

 

canidia Sparrman, 1768  (Papilio). Amoenula Academiae, 7: 504. TL: China . Had been originally described as a good species.

·       canidia Sparrman, 1768 (Papilio). D: South Ussuri district.

·       palaearctica Staudinger, 1886  (Pieris). Stettiner entomologische Zeitschrift, 47: 198. TL: “Südfergana, Ostbuchara”. Had been originally described as a subspecies of canidia. D: All mountains of Middle Asia ex. Pamir .

 

dulcinea Butler , 1882  (Ganoris). Annales of the Magazin of the natural History, (5) 9: 18 – 19. TL: Dabeyde Bay , southern Wusuli river. Had been originally described as a good species.

·       dulcinea Butler , 1882 (Ganoris). D: Amur district; Ussuri district.

·       saghalensis Nakahara, 1926  (Pieris). Insecutor Inscit. menestr., Washington , 14: 46. TL: “Saghalien”. Had been originally described as a subspecies of melete. D: Sakhalin .

 

ergane Geyer  in Hübner, [1827 – 1828] (Papilio). Sammlung europäischer Schmetterlinge, Augsburg : Tab. 184; Fig. 904 – 907. TL: Sicily , Ragusa (Hemming, 1937, 1: 220). Had been originally described as a good species.

·       detersa Verity, [1908]  (Pieris). Rhopalocera palaearctica, Firenze , [1]: 153; pl. 33, fig. 2. TL: “Berud Dagh, monts Taurus”. Had been originally described as a variety of ergane. D: Little Caucasus ; Transcaucasus.

 

euorientis Verity, [1908]  (Pieris). Rhopalocera palaearctica, Firenze , [1]: 147, 167; pl. 32, fig. 44; pl. 49, fig. 12. TL: Sajan, “Muorku”. Had been originally described as a subspecies of napi.

Remark. The specific level was given by V.Lukhtanov & A.Lukhtanov (1994: 62): “In Südsibirien (im Sajan-Gebirge, Tannu-Ola-Gebirge und Transbaikal-Gebiet) kommt auch die sehr ähnliche [zur banghaasi] Art P. euorientis... vor, die für den Dshungarischen Alatau nachgewiesen wurde (Eitschberger, [1984])”.

·       euorientis Verity, [1908] (Pieris). D: Sayan; Tuva; Praebaikal; Transbaikal; ?Dzhungaria.

·       sauron Yakovlev, 2004 (Pieris). Helios, Moscow , 5: 233 – 234; pl. 16, figs. 1 – 8. TL: “East Kazakhstan, Saur Mts., Chagan-Obo, h 1600 – 1800 m , 47º 02’ N; 84º 54’ E”. Had been originally described as a subspecies of euorientis. D: Saur Mts.

 

krueperi Staudinger, 1860  (Pieris). Wiener entomologische Monatsschrift, 4: 19. TL: Greece . Had been originally described as a good species.

·       devta de Nicéville, 1883  (Mancipium). Journal of the Asiatic Society, Bengal , 1883: 82. TL: Ladak. Had been originally described as a good species. D: Mountains of Middle Asia .

 

mahometana Grum-Grshimailo, 1888  (Pieris). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 22: 304. TL: “Darwaz”. Had been originally described as a variety of krueperi.

Remark. The specific status by Wyatt & Omoto (1966: 148).

·       mahometana Grum-Grshimailo, 1888 (Pieris). D: Darvaz; West Pamir .

 

manii Mayer, 1851  (Pontia). Stettiner entomologische Zeitschrift, 12: 151. TL: Split , Dalmatia , Yugoslavia . Had been originally described as a good species.

·       manii Mayer, 1851 (Pontia). D: Southern part of Transcaucasus.

 

melete Ménétriès, 1858  (Pieris). Enumeratio corporum animalium Musei imperialis academiae scientarum Petropolitanae, Petropoli, 11: 72. TL: Japan . Had been originally described as a good species.

·       melete Ménétriès, 1858 (Pieris). D: Ussuri district; Amur district (from Blagovestshensk to Gorin river).

 

napi Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 468. TL: Sweden . Had been originally described as a good species.

·       balcarica Wojtusiak & Niesiołowski, 1947  (Pieris). Prace Muzei przyrodne polone Akademie Umiejetne, Praha, 6: 45. TL: “area between Naltchik and Kara-su... from 1000 – 1500 m ”. Had been originally described as a subspecies of napi. D: Great Caucasus; Small Caucasus .

·       dubiosa Röber  in Seitz, [1907] (Pieris). Die Groβschmetterlinge der Erde. Die Groβschmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart , 1: 46; Taf. 20 e. TL: “Andalusien, Klein-Asien”. Had been originally described as a forma of napi. D: Transcaucasus.

·       muchei Eitschberger, [1984]  (Pieris). Herbipoliana, Marktleuthen, 1: 135; Taf. 411, Abbn. 19/20. TL: Kazakhstan , Transilian Alatau, Almaarasan. Had been originally described as a subspecies of napi. D: All Middle Asiatic mountains without Pamir and Transalai Mts. ; Dzhungaria; Tarbagatai.

·       napi Linnaeus, 1758 (Papilio). D: European part; Crimea ; South Ural.

 

elbursina Bytinski-Salz, 1937  (Pieris). Entomologist’s Record and Journal of Variation, London , Suppl. 49: 1. TL: Iran , Elburs. Had been originally described as a subspecies of napi.

·          elbursina Bytinski-Salz, 1937 (Pieris). D: Talysh.

 

nesis Fruhstorfer, 1909  (Pieris). Internaionale Entomologische Zeitschrift, Frankfurt am Main, 3 (16): 88. TL: Japan , Hokkaido , Sapporo . Had been originally described as a subspecies of napi.

·       nesis Fruhstorfer, 1909 (Pieris). D: Sakhalin; South Kuriles .

 

ochsenheimeri Staudinger, 1886  (Pieris). Stettiner entomologische Zeitschrift, 47: 199. TL: “ Namangan ”. Had been originally described as a good species.

·       gerhardi Eitschberger, [1984]  (Pieris). Herbipoliana, Marktleuthen, 1: 228. TL: “ USSR , 3000 m , Geb. Peter d. Grossen, Ganischou”. Had been originally described as a subspecies of ochsenheimeri. D: Darvaz; Gissar.

·       ochsenheimeri Staudinger, 1886 (Pieris). D: Alai; Inner Tian Shan .

 

persis Verity, 1922  (Pieris). Entomological Record and Journal of Variation, London , 34: 140. Had been originally described as a race of napi.

·       persis Verity, 1922 (Pieris). D: West Kopet Dagh: upper course of Sajvan, Evshanli; Central Kopet Dagh: Deshikli-dere valley, Germab, Nizhnie Chuli (Dubatolov et al., 1989).

 

pseudorapae Bowden & Riley, 1967  (Pieris). Redia, 50: 379. TL: Syria , Beyrouth – by the lectotype (Bowden & Riley, 1967). Had been established in the quadrinominal combination Pieris napi napi pseudorapae; the new rank by Bowden & Riley, 1967.

·       suffusa Verity, [1908]  (Pieris). Rhopalocera palaearctica, Firenze , [1]: 143; pl. 32, fig. 21, 22. TL: “Transcaucasie, Jelizavetopol”. Had been originally described as a forma of napi. D: Caucasus; Transcaucasus; Kopet Dagh; Crimea .

 

rapae Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 468. TL: Sweden . Had been originally described as a good species.

·       debilis Alphéraky, 1889  (Pieris). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 70 - 71. TL: Lob Nor. Had been originally described as a variety of rapae.  D: Middle Asia ; Dzhungaria.

·       rapae Linnaeus, 1758 (Papilio). D: European part; Siberia; Far East .

 

tomariana Matsumura, 1929  (Pieris). Insecta Matsumurana, Sapporo , 2 (4): 191, 197. TL: Kunashir, Tomari. Had been originally described as a subspecies of napi.

·       tomariana Matsumura, 1929 (Pieris). D: South Kuriles .

 

transiens Verity, [1908]  (Pieris). Rhopalocera palaearctica, Firenze , [1]: 138. TL: Askold Island . Had been originally established in the quadrinominal combination Pieris melete melete transiens.

·       transiens Verity, [1908] (Pieris). D: Askold Island .

 

Magazin für Insektenkunde, 6: 283. TS: Papilio daplidice Linnaeus, 1758.

 

callidice Hübner, [1800]  (Papilio). Sammlung europäischer Schmetterlinge, Augsburg , 1: Fig. 408, 409, 522. TL: Alps . Had been originally described as a good species.

·       callidice Hübner, [1800] (Papilio). D: Carpathians; Great Caucasus; Small Caucasus; Altai; Sayan; Praebaikal; Tuva; Transcaucasus.

·       daritshevae Dubatolov in Tuzov, 1993  (nomen nudum) (Pontia). The synonymic list of the butterflies from the ex-USSR, Moscow : 20. TL: “Kopet-Dagh, S.Gissar”. D: TL.

·       halasia Huang & Murayama, 1992  (Pontia). Tyô to Ga, Tokyo , 43: 3; fig. 12. TL: “Halasi, Altai, 1400 – 1700 m , grassland in forest zone”. Had been originally described as a subspecies of callidice. D: Ukok in SW Altai.

·       hinducucica Verity, [1911]  (Pieris). Rhopalocera palaearctica, Firenze , [1]: 326; pl. 52, fig. 13. TL: Hindukush. Had been originally described as a race of callidice. D: West Pamir .

·       kalora Moore , 1865  (Pieris). Proceedings of the zoological Society of London, 1865: 489 – 490; pl. 31, fig. 15. TL: “Passes over the spurs of the Himalayas which run from Spiti southward to the Sutlej ”. Had been originally described as a good species. D: Tian Shan, Gissar (without type locality of daritshevae), Alai, Darvaz, Pamir .

·       nelsoni Edwards, 1883  (Pontia). The Canadian Entomologist, Ottawa , 15: 324. TL: St. Michael’s Alaska . Had been originally described as a variety of callidice. D: Polar tundra of Siberia; Far East; Kamchatka ; Chukotka.

Remark. Y.P.Korshunov (1998: 13) wrote: “В книге московских коллекционеров "Butterflies of Russia", v. 1, 1997 значится, что callidice и американский occidentalis конспецифичны, и похоже только на этом основании маккинлианский подвид nelsoni с Аляски выдают за жителя тундры Сибири, Дальнего Востока, Камчатки и Чукотки. Предполагать, конечно, можно всякое, но известны специальные опыты и вывод о генетической несовместимости конкретно между P. callidice и P. occidentalis nelsoni (W.H.Edwards, 1883) – смотри работы А.М.Шапиро (1975 et al.). А вообще и бабочки под номерами 10 – 12 на таблице 29 не совпадают с изображением nelsoni на таблице 97 (No. 29) в книге W.H.Howe (1975)”. [In the book of the Moscow collectors " Butterflies of Russia ", v. 1, 1997 appears, that callidice and American occidentalis are conspecific, and is similar only on this basis the McCinley subspecies nelsoni from Alaska give out for the inhabitant of tundra of Siberia, Far East, Kamchatka and Chukotka. To assume, certainly, everyone is possible, but the special experiences and conclusion about genetic incompatibility particularly between P. callidice and P. occidentalis nelsoni (W.H.Edwards, 1883) – look works of А.М.Shapiro (1975 et al.). And in general and the butterflies under numbers 10 – 12 on the plate 29 do not coincide with the image nelsoni on the plate 97 (No. 29) in the book W.H.Howe (1975)]. I believe, what not superfluous will result here following information. According to work Gary M. Marrone (1994: 233, 240), the nominotypical subspecies of  Pontia occidentalis in USA lives in state South Dakota . The subspecies nelsoni was described from the McKinley mountains  in Alaska , and differs from occidentalis so strongly, that it could be accepted for the specific level, if not the comparison it with callidice. As is shown by such comparison, nelsoni there is not that other, as the subspecies of circumpolar callidice, here again opinion of Y.P.Korshunov in a root is incorrect.

 

chloridice Hübner, [1813]  (Papilio). Sammlung europäischer Schmetterlinge, Augsburg , 1: Tab. 141, Fig. 712 – 715. TL: Europa. Had been originally described as a good species.

·       chloridice Hübner, [1813] (Papilio). D: European part northwards to N.Novgorod region; Siberia; Caucasus; Transcaucasus; Kazakhstan; Middle Asia; Dzhungaria.

 

daplidice Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 468. TL: North-West Africa (Wagener, 1988: 30). Had been originally described as a good species.

·       ssp? D: Correctly known only from southern Ussuri district (Kesküla, 1993: fig. 1).

 

edusa Fabricius, [1777]  (Papilio). Genera Insectorum..., Chilonii: 225. TL: Kiel . Had been originally described as a good species.

·       edusa Fabricius, [1777] (Papilio). D: European part northwards to Estonia (Kesküla, 1990); Caucasus; Siberia; Amur district; Ussuri district; Crimea; Transbaikal; Sakhalin.

·       kulyginskyi Korshunov, 1998  ([Pontia]). Novye opisanija i utochnenija dlja knigi "Dnevnye babochki asiatskoj chasti Rossii", Novosibirsk : 12. TL: "Бреды, степь" [Bredy, steppe, South Ural]. Had been originally described as a  subspecies of daplidice. D: South Ural.

Remark. In the present work I leave the status kulyginskyi on conscience of the author of the original description, having changed only combination of this taxon – from daplidice in edusa. The business that daplidice does not live on South Ural, to what the publications of numerous western and domestic authors testify. I think also, that the author of kulyginskyi has described an aberratio edusa, however to check up it while I can not because of absence of a material.

·       persica Bienert, 1869  (Pieris). Lepidopterologische Ergebnisse der Reise im Persien: 26. TL: “Pers. mer.” – by the lectotype (Kesküla, 1993: 3). Had been originally described as a variety of daplidice. D: Kopet Dagh; Transcaucasus.

 

glauconome Klug, 1829  (Pontia). Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustrate redierunt, Berolini: e; Tab. 3. TL: North Syria . Had been originally described as a good species.

·       glauconome Klug, 1829 D: Kopet Dagh; South Gissar .

Annales of the Magazin of the natural history, (5) 1: 228. TS: Mesapia shawii Bates in Henderson & Hume, 1873.

 

nekrutenkoi Kreuzberg, 1989 , stat. n. (Baltia). Vestnik zoologii, Kiev , 1989 (6): 39 – 41; ris. 9. TL: “Центральный Тянь-Шань, Корум-Кечу” [ Central Tian Shan , Korum-Kechu]. Had been originally described as a subspecies of butleri.

·       nekrutenkoi Kreuzberg, 1989 (Baltia). D: Inner Tian Shan .

 

shawii Bates  in Henderson & Hume, 1873 (Mesapia). Lahore to Yarkand. Incidents of the route and natural history of the countries traversed by the expeditions of 1870, London : 305. TL: “on the Chang Lang Pass : alt. 18,000 ft .”. Had been originally described as a good species.

·       wyatti Schulte, 1989  (Baltia). Nachrichten des entomologischen Vereins Apollo, N.F., Frankfurt am Main, 10: 296; Taf. 1, Fign. 11, 12. TL: “ Afghanistan , Anjuman-Tal, 2800 – 4250 m , Hindukusch”. Had been originally described as a subspecies of shawi [sic!]. D: Pamir .