Family Nymphalidae Swainson, 1827

Subfamily Apaturinae Boisduval, 1840

 

Proceedings of the Zoological Society of London, 1822: 240. TS: Limenitis dichroa Kollar, [1844].

 

princeps Fixsen, 1887 (Apatura). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 289, pl. 13, fig. 7. TL: "Pung-Tung" in Corea. Had been originally described as a good species.

·          princeps Fixsen, 1887 (Apatura). D: South Ussuri region; southern part of Khabarovsk Territory.

 

Magazin für Insektenkunde, 6: 280. TS: Papilio iris Linnaeus, 1758.

 

ilia [Denis & Schiffermüller], [1775] (Papilio). Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend, Wien: 172. TL: Vienna. Had been originally described as a good species.

·          ilia [Denis & Schiffermüller], [1775] (Papilio). D: European part; Siberia (without northern part); Transbaikal; Caucasus; Sayan; Carpathians; North Kazakhstan.

·          krylovi Kurentzov, 1937 (Apatura). Vestnik dalnevostochnogo filiala Akademii Nauk SSSR, Vladivostok, 26: 116. TL: "северныйСихотэ-Алинь" [Northern Sikhote Alin]. Had been originally described as a subspecies of ilia. D: North Sikhote Alin; Middle Sikhote Alin; Sakhalin; South Kuriles.

·          ussuriensis Kurentzov, 1937  (Apatura). Vestnik dalnevostochnogo filiala Akademii Nauk SSSR, Vladivostok, 26: 116. TL: "Приморскийкрай, Уссурийскийзаповедник" [Ussuri region; Ussuri reservation] – by the syntypes (Azarova, 1986: 125). Had been originally described as a subspecies of ilia. D: Amur region; Ussuri region.

 

iris Linnaeus, 1758 (Papilio). Systema Naturae… (ed. 10), 1: 476. TL: Germany, England. Had been originally described as a good species.

·          abramovi Kurentzov, 1970 (Apatura). Bulavousye cheshuekrylye Dalnego Vostoka SSSR, LeningraD: 88; ris. 81 (2). TL: "внижнемтечениир. Амура (р. Горин) инапобережье (р. Самарга)" [in the under course of Amur river (river Gorin) and in the coast (river Samarga)). Had been originally described as a subspecies of iris. D: Amur region; Ussuri region.

·          iris Linnaeus, 1758 (Papilio). D: Central and southern areas of European part; South Siberia; West Siberia: Omsk region (Knyazev & Kosterin, 2003); Transbaikal; North Kazakhstan; Carpathians.

 

metis Freyer, 1829 (Apatura). Beiträge zur Geschichte der europäischen Schmetterlinge, Augsburg, 2: 67; Tab. 67, Abb. 1. TL: “Syrmien” [Dunav river in Croatia]. Had been originally described as a good species.

·          bunea Herrich-Schäffer, [1845] (Apatura). Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu Jakob Hübner's Sammlung europäischer Schmetterlinge. Umrisstafeln mit Erklärung. Nachträge. Systema Lepidopterorum. Index alphabetico-synonymicus ad Vol. I – V. Index universalis specierum et generum, Regensburg, 1: 45; Fig. 161 – 164. TL: "aus dem Caucasus". Had been originally described as a variety of ilia. D: Caucasus; Transcaucasus; South of European part; South Ural; North Kazakhstan.

·          doii Matsumura, 1928 (Apatura). Insecta Matsumurana, Sapporo, 2 (4): 191, 199. TL: Kunashir. Had been originally described as a subspecies of iris. D: South Kuriles; Sakhalin.

·          irtyshika Korshunov, 1982 (Apatura). Gelminty, klestshi i nasekomye, Novosibirsk: 88. TL: "Павлодарскаяобл., с. Качиры" [Pavlodar region, Kachiry]. Had been originally described as a subspecies of metis. D: North-East Kazakhstan; South Siberia to Baikal Lake.

·          separata Tuzov, 1999 (Apatura). Helios, Moscow, 1: 104 - 105. TL: “Chita Reg., Talacha loc., Shilka distr.”. Had been originally described as a subspecies of metis. D: Transbaikal.

·          substituta Butler, 1873 (Apatura). Cistula Entomologica, 1: 159. TL: Japan. Had been originally described as a good species. D: Amur region; Ussuri region.

 

Lepidoptera Indica, Madras, 3: 10. TS: Atyma nycteis Ménétriès, 1859.

 

nycteis Ménétriès, 1859 (Athyma). Bulletin de l’Academie des Sciences, Sankt-Petersburg, 17: 215. TL: "Amur". Had been originally described as a good species.

·          nycteis Ménétriès, 1859 (Athyma). D: Amur region; Ussuri region.

[nom. nov. pro Bremeria Moore, 1896, nec. Bremeria Alphéraky, 1892]. Chlenistonogie i gelminty, Novosibirsk: 52. TS: Adolias schrenckii Ménétriès, 1859.

 

schrenckii Ménétriès, 1859 (Adolias). Bulletin de l’Academie des Sciences, Sankt-Petersburg, 17: 215. TL: “montagnes du Chingan”. Had been originally described as a good species.

·          schrenckii Ménétriès, 1859 (Adolias). D: Amur region; Ussuri region; South Transbaikal.

 

TS: Apatura morgiana Westwood in Doubleday, [1850].

 

fenestra Leech, 1891 (Apatura). Entomologist, London, 24, Suppl.: 26. TL: West China: “Omeishan”. Had been originally described as a good species.

·       fenestra Leech, 1891 (Apatura). D: Southern part of Ussuri region.

Catalog der Lepidopteren des europäischen Faunengebietes, Berlin (Aufl 1): 17. TS: Nymphalis jonia Fischer de Waldheim & Eversmann, 1851.

 

jonia Fischer de Waldheim & Eversmann, 1851 (Nymphalis). Entomographia imperii Rossici, Moscow, 5: 111; Tab. 13, fig. 1, 2. TL: Transcaucasus. Had been originally described as a good species.

·          jonia Fischer de Waldheim & Eversmann, 1851. D: Transcaucasus.

 

Subfamily Charaxinae Leech, 1890

 

Die Schmetterlinge von Europa, Leipzig, 4: 18. TS: Papilio jasius Linnaeus, 1767.

 

jasius Linnaeus, 1767 (Papilio). Systema Naturae… (ed. 12), 1 (2): Errata. TL: Europa. D: Only some findings are known from Crimea.

 

Subfamily Limenitinae Butler, 1869

 

Transactions of the Kansai entomological Society, 23: 34. TS: Limenitis pratti Leech, 1890.

 

pratti Leech, 1890 (Limenitis). Entomologist, London, 23: 34. TL: Central China, “Chang-Yang”. Had been originally described as a good species.

·          eximia Moltrecht, 1909 (Limenitis). Entomologische Zeitschrift, Frankfurt am Main, 22: 184. TL: "in Okeanskaja, Villenort, 25 km von Wladiwostok am bewaldeten Meersstrande,… am Westabhang der Sichota-Alin-Bergskette", "120 km östlich von Nikolsk-Ussurijsk". D: Ussuri region.

 

Magazin für Insektenkunde, 6: 281. TS: Papilio populi Linnaeus, 1758.

 

amphissa Ménétriès, 1859 (Limenitis). Bulletin de l’Academie des Sciences, Sankt-Petersburg, 17: 215. TL: “des monts Boureia, et près de l’embourchure de l’Oussouri”. Had been originally described as a good species.

·          amphissa Ménétriès, 1859 (Limenitis). D: Ussuri region; Amur region from Zea to Gorin river.

 

camilla Linné, 1764 (Papilio). Museum S:ae R:ae M:tis Ludovicae Ulricae Reginae Svecorum, Gothorum, Vandalorumque etc.etc.etc. In quo animalia rariora, exotica, imprimis Insecta et Conchilida describuntur et determinatur, prodromi instar editum, Holmiae: 304. TL: Germany. Had been originally described as a good species.

·          camilla Linné, 1764 (Papilio). D: European part; Caucasus; Transcaucasus.

·          japonica Ménétriès, 1857 (Limenitis Enumeratio corporum animalium musei imperialis academie scientarum Petropolotanae, Petropoli, 2: 93. TL: “Japon”. Had been originally described as a variety of sibylla. D: Sakhalin; Amur region; Ussuri region.

 

doerriesi Staudinger, 1892 (Limenitis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 173; Tab. 14, fig. 1. TL: “Suschan-Gebiete”. Had been originally described as a good species.

·          doerriesi Staudinger, 1892 (Limenitis). D: Ussuri region.

 

helmanni Kindermann in Lederer, 1853 (Limenitis). Verhandlungen des zoologisch-botanischen Vereins, Wien, 3: 356; Taf. 1, Fig. 4. TL: Altai, “Usbucharminsk”. Had been originally described as a good species.

·          duplicata Staudinger, 1892 (Limenitis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 6: 171. TL: “Amur”. Had been originally described as a variety of helmanni. D: Ussuri region.

·          helmanni Kindermann in Lederer, 1853 (Limenitis). D: Altai; Kusnetzky Alatau.

·          montana Kurentzov, 1941 (Limenitis). Trudy gorno-tajozhnoj stanzii Akademii Nauk SSSR, 4: 363. TL: "Приморскийкрай, СреднийСихотэ-Алинь" [Ussuri region, Middle Sikhote Alin]. Had been originally described as a subspecies of helmanni. D: East Transbaikal; Amur region; northern Ussuri region.

 

homeyeri Tancré, 1881 (Limenitis). Entomologische Nachträge, 8: 120. TL: "Blagowesschensk; Raddefskaja". Had been originally described as a good species.

·          homeyeri Tancré, 1881 (Limenitis). D: Ussuri region.

 

lepechini Erschoff in Fedtschenko, 1874 (Limenitis). Reise nach Turkestan, Sankt-Petersburg, 2 (5) (7): 47. TL: "...circa Maracanda (2150’ sm.), volat junio mense". Had been originally described as a good species. D: Gissar; Darvaz.

·          lepechini Erschoff in Fedtschenko, 1874 (Limenitis). D: Gissar; Darvaz.

 

moltrechti Kardakov, 1928 (Limenitis). Entomologische Mitteilungen, Berlin, 17: 269. TL: “Narva”, Ussuri region. Had been originally described as a good species.

·          moltrechti Kardakov, 1928 (Limenitis). D: Middle Amur region; Ussuri region.

 

populi Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 476. TL: Sweden. Had been originally described as a good species.

·          fruhstorferi Krulikowsky, 1909 (Limenitis). Revue Russe d'Entomologie, Moscow, 9: 111. TL: "Rossia orientali". Had been originally described as a subspecies of populi. D: South-East of European part; Ural; West Siberia.

·          halasiensis Huang & Murayama, 1992 (Limenitis). Tyô to Ga, Tokyo, 43 (1): 9; fig. 21, 22. TL: "Halasi, Altai, 1400 – 1600 m, grassland in forest zone". Had been originally described as a subspecies of populi. D: SW Altai.

·          populi Linnaeus, 1758 (Papilio). D: European part; Caucasus; North Kazakhstan.

·          ussuriensis Staudinger, 1887 (Limenitis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 143. TL: “Ussuri”. Had been originally described as a variety of populi. D: Amur region; Ussuri region; Transbaikal.

 

reducta Saudinger, 1901 (Limenitis). Catalog der Lepidopteren des palaearktischen Faunengebietes, Berlin (Aufl. 1): 22. TL: “Arm.[enia] or.[ientalis]; Hyrc.[ania]”. Had been originally described as a variety of camilla.

·          reducta Staudinger, 1901 (Limenitis). D: Southern part of European Russia, Ukraina; Caucasus, Transcaucasus.

 

sydyi Kindermann in Lederer, 1853 (Limenitis). Verhandlungen des zoologisch-botanischen Vereins, Wien, 3: 357; Taf. 1, Fig. 3. TL: “Ustbuchtarminsk”. Had been originally described as a good species.

·          latefasciata Ménétriès in Schrenck, 1859 (Limenitis). Reisen und Forschungen im Amur-Lande, Sankt-Petersburg, 2 (1): 29. TL: “Pakhale”. Had been originally described as a variety of sydyi. D: South of Far East.

·          sydyi Kindermann in Lederer, 1853 (Limenitis). D: Ust-Kamenogorsk and Zmeinogorsk vicinity; East Transbaikal; Amur region; Kalbinsky Mts.

 

Lepidoptera Indica, 3: 46. TS: Diadema raddei Bremer, 1861.

 

raddei Bremer, 1861 (Diadema). Bulletin de l’Academie des Sciences, Sankt-Petersburg, 3: 547. TL: “Bureja-Gebirge” . Had been originally described as a good species.

·          raddei Bremer, 1861 (Diadema). D: Ussuri region; Amur region.

 

Magazin für Insektenkunde, 6: 282. TS: Papilio aceris Esper, [1783]

 

alwina Bremer & Grey in Motshulsky, 1852 (Neptis). Études d'Entomologie, Helsingfors, 6: 60. TL: China near Peking. Had been originally described as a good species.

·          alwina Bremer & Grey in Motshulsky, 1852 (Neptis). D: Amur region in vicinities of towns Khabarovsk and Komsomolsk-na-Amur; Ussuri region.

 

andetria Fruhstorfer in Seitz, 1912 (Neptis). Die Gross-Schmetterlinge der Erde, Stuttgart, 9: 609. TL: Vladivostok. Had been originally described as a subspecies of pryeri. The specific status by P.Y.Gorbunov (2001: 161).

·       andetria Fruhstorfer in Seitz, 1912 (Neptis). D: Amur region, eastern part; southern Ussuri region.

 

ilos Fruhstorfer, 1909 (Neptis). Entomologische Zeitschrift, Frankfurt am Main, 23: 42. TL: “?Amur Gebiet”. Had been originally described as a subspecies of themis.

·       ilos Fruhstorfer, 1909 (Neptis). D: Southern Ussuri region.

 

kusnetzovi Kurentzov, 1949 (Neptis). Revue d'Entomologie de l'URSS, Léningrad, 30 (3 – 4): 362; ris. 1, 2. TL: "р. Колумбе" [Kolumbe river]. Had been originally described as a good species.

·          kusnetzovi Kurentzov, 1949 (Neptis). D: Middle Sikhote-Alin.

 

philyra Ménétriès, 1859 (Neptis). Bulletin de l’Academiae des Sciences, Sankt-Petersburg, 17: 214. TL: "...environs de Marienpost". Had been originally described as a good species.

·          philyra Ménétriès, 1859 (Neptis). D: Ussuri region; Amur region.

 

philyroides Staudinger, 1887 (Neptis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 146. TL: “Raddeefka”. Had been originally described as a good species.

·          philyroides Staudinger, 1887 (Neptis). D: Amur region; Ussuri region.

 

pryeri Butler, 1871 (Neptis). Transactions of the Royal entomological Society of London, 1871: 403. TL: “Shanghai“ in East China. Had been originally described as a good species.

·          pryeri Butler, 1871 (Neptis). D: Southern border of Ussuri area (Yuzhnoe Primor’ye [ЮжноеПриморье]). This species in the book of P.Y.Gorbunov (2001: 161) listed as very possible species for the Russian fauna; in the collections of insects of the Zoological Museum of the Nizhny Novgorod State University are two specimens of this species from the collecting trips of Dr G.A.Anufriev in the Southern Ussuri region (Partizansk vicinity).

 

rivularis Scopoli, 1763 (Papilio). Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietates. Methodo Linneana, Vindobonae: 165; Fig. 443. TL: Carinthia. Had been originally described as a good species.

·          ludmilla Herrich-Schäffer, 1856 (Neptis). Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu Jakob Hübner's Sammlung europäischer Schmetterlinge. Umrisstafeln mit Erklärung. Nachträge. Systema Lepidopterorum. Index alphabetico-synonymicus ad Vol. I – V. Index universalis specierum et generum, 6: 1; Fign. 301, 302. TL: South Russia. Had been originally described as a variety of rivularis. D: Transcaucasus; Caucasus; deserts of Middle Asia and Kazakhstan; Tian Shan; Darvaz; Gissar; Alai; Crimea; Carpathians; European part; Ural.

·          magnata Heyne in Rühl, [1895] (Neptis). Die palaearktische Grossschmetterlinge und ihre Naturgeschichte, Leipzig, 1: 776. TL: “Kentei”. Had been originally described as a variety of aceris. D: South Siberian mountains; Amur region; Ussuri region; Sakhalin; South Kuril Islands.

 

sappho Pallas, 1771 (Papilio). Reisen durch verschieden Provinzen des Russischen Reichs in den Jahren 1768 – 1774, Sankt-Petersburg, 1: 471. TL: Volga. Had been originally described as a good species.

·          sappho Pallas, 1771 (Papilio). D: European part; Siberia (without north); Kazakhstan (north); Ural.

 

speyeri Staudinger, 1887 (Neptis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 145; tab. 7, fig. 3. TL: “Ussuri”. Had been originally described as a good species.

·          speyeri Staudinger, 1887 (Neptis). D: Ussuri region – to Kievka river in north and to Khabarovsk by the Ussuri river valley.

 

thisbe Ménétriès, 1859 (Neptis). Bulletin de l’Academie des Sciences, Sankt-Petersburg, 17: 214. TL: “des monts Boureïa, ...et pres de l’Oussouri”. Had been originally described as a good species.

·          thisbe Ménétriès, 1859 (Neptis). D: Middle Amur region; South Transbaikal.

·          ussuriensis Kurentzov, 1970 (Neptis). Bulavousye cheshuekrylye Dalnego Vostoka SSSR, LeningraD: 92 – 93. TL: "ВБуреинскихгорахиюжнеевбвссейнеУссури" [in the Bureinskye mountains and souther in the Ussuri river basin]. Had been originally described as a subspecies of thisbe. D: Ussuri region.

 

tshetvericovi Kurentzov, 1936 (Neptis). Lambillionea, Bruxelles, [36] (7): 185 – 190; ris. 3. TL: "Ussuri". Had been originally described as a good species.

·          tshetvericovi Kurentzov, 1936 (Neptis). D: Transbaikal in the rivers' valleys of Shilka and Amgun' as well as in the Erman mountain ridge; Amur region; Ussuri region.

Remark. The specimens of this very interest species which collected in Transbaikal differ from the Ussurian butterflies of tshetvericovi by some external charakters: size, submarginal pattern and ground colour. It is possible that Transbaicalian population of tshetvericovi is a good subspecies.

 

Subfamily Nymphalinae Swainson, 1827

Verzeichniss bekannter Schmetterlinge, Augsburg, 3: 36. TS: Papilio c-album Linnaeus, 1758.

 

c-album Linnaeus, 1758 (Papilio). Systema Naturae… (ed. 10), 1: 477. TL: [Sweden]. Had been originally described as a good species.

·          c-album Linnaeus, 1758 (Papilio). D: European part; Caucasus; Ural; Carpathians; Crimea; Kazakhstan (without south); Siberia; Far East (without south).

·          hamigera Butler, 1877 (Vanessa). Annales of the Magazin of Natural History, 19: 92. TL: “about 370 miles from Tokey”. Had been originally described as a good species. D: Amur region; Ussuri region.

·          interposita Staudinger, 1881 (Polygonia). Stettiner entomologische Zeitschrift, 42: 286. TL: "Ala Tau". Had been originally described as a variety of c-album. D: Middle Asiatic mountains.

 

c-aureum Linné, 1767 (Papilio). Systema Naturae… (ed. 12), 1: 477. TL: “Asia”. Had been originally described as a good species.

·          c-aureum Linné, 1767 (Papilio). D: Ussuri region; Amur region.

 

egea Cramer, [1775] (Papilio). Papillons Exotiques: pl. 78, figs. C, D. TL: “Constantinopel”, “Smirna”. Had been originally described as a good species.

·            egea Cramer, [1775] (Papilio). D: Caucasus, Transcaucasus, Southern part of Russia, Kopeth-Dagh.

 

undina Grum-Grshimaïlo, 1890 (Vanessa). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 424; pl. 17, fig. 1. TL: "Oche". Had been originally described as a good species.

·          undina Grum-Grshimaïlo, 1890 (Vanessa). D: Middle Asiatic mountains.

 

Dnevnye babochki asiatskoj chasti Rossii, Ekaterinburg: 81. TS: Papilio l-album Esper, [1780].

 

vau-album [Denis & Schiffermüller], [1775] (Papilio). Ankündung eines systematischen Werkes von der Schmetterlinge der Wienergegend, Wien: 176. TL: Vienna. Had been originally described as a good species.

·          arbustus Churkin & Zhdanko, 2002 (Nymphalis). Helios, Moscow, 3: 129 – 130. TL: “Kirghizstan, Alai Mts., Dugoba River, 2600 m”. Had been originally described as a subspecies of vaualbum. D: Middle Asiatic mountain.

·          samurai Fruhstorfer, 1907 (Vanessa). Entomologische Zeitschrift, Frankfurt am Main, 21: 23. TL: Ussuri. Had been originally described as a variety of vau-album. D: Amur region; Ussuri region; Sakhalin.

·          vau-album [Denis & Schiffermüller], [1775] (Papilio). D: European part; Siberia (without north); Caucasus; Kazakhstan; Carpathians.

 

Zwierząt domowych i dzikich, osobliwie kraiowych, historyi naturalney początki i gospodarstwo. Potrzebnych i potecznych domowych chowanie, rozmnożenie, chorób leczenie, dzikich łowienie, oswoienie, zażycie, szkodiwych zas wygubienie. O owadzie i robakach. Warszawa, 4: 86. TS: Papilio polychloros Linnaeus, 1758.

 

polychloros Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 477. TL: Sweden. Had been originally described as a good species.

·          polychloros Linnaeus, 1758 (Papilio). D: Middle and Southern part of European ex-USSR; Caucasus; Transcaucasus; Ural; Crimea; South Siberian mountains; Kopet Dagh; Tian Shan; Gissar.

 

xanthomelas [Denis & Schiffermüller], [1775] (Papilio). Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend, Wien: 175. TL: "Aus dem deutschen Reich und aus der Steyermark". Had been originally described as a good species.

·          hazara Wyatt & Omoto, 1966 (Nymphalis). Entomops, Nicé, 5: 153; figs. TL: "near Panjao, W. Koh-i-Baba Mts., 2,700 m". Had been originally described as a subspecies of xanthomelas. D: Alai; Pamir; Darvaz.

·          sachalinensis Matsumura, 1925 (Vanessa). Journal of the College of Agriculture of Hokkaido  Imperial University of Sapporo, 15 (3): 197. TL: Sakhalin: “Nairo, Tim, Arcovo”. Had been originally described as a subspecies of xanthomelas. D: Sakhalin; Kuriles.

·          xanthomelas [Denis & Schiffermüller], [1775] (Papilio). D: European part; Siberia; Far East; Caucasus; Kazakhstan; Crimea; Tian Shan; Gissar; Kopet Dagh.

 

TS: Papilio antiopa Linnaeus, 1758.

 

 

 

antiopa  Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 476. TL: “...etiam en Americae”. Had been originally described as a good species.

·          antiopa Linnaeus, 1758 (Papilio). D: European part; Siberia; Middle Asia (without Pamir and Alai); Caucasus.

·          asopos Fruhstorfer, 1909 (Vanessa).  Internationale Entomologische Zeitschrift, Guben, 3: 94. TL: Japan. Had been originally described as a subspecies of antiopa. D: Sakhalin.

·          borealis Wnukowsky, 1927 (Vanessa). Mitteilungen der Münchener entomologischen Gesellschaft, 17: 69.  TL: Yakutsk. Had been originally described as a subspecies of antiopa. D: Far East.

 

Magazin für Insektenkunde, 6: 281. TS: Papilio atalanta Linnaeus, 1758.

 

atalanta Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 478. TL: Sweden. Had been originally described as a good species.

·          atalanta Linnaeus, 1758 (Papilio). D: European part; Ural; West Siberia; Altai; Sayan; Carpathians; Crimea; Caucasus; North Tian Shan; Dzhungaria; Kazakhstan.

 

indica Herbst et Jablonsky, 1794 (Papilio). Natursystematik der Insekten, Berlin, 7: 171; Taf. 180, Fig. 1, 2. TL: India. Had been originally described as a variety of atalanta.

·          indica Herbst et Jablonsky, 1794 (Papilio). D: Amur region; Ussuri region; Sakhalin; possible in South Kuriles; Transbaikal.

 

Magazin für Insektenkunde, 6: 281. TS: Papilio cardui Linnaeus, 1758.

 

cardui Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 475. TL: Sweden. Had been originally described as a good species.

·          cardui Linnaeus, 1758 (Papilio). D: All territory of the ex-USSR.

 

Verzeichniss bekannter Schmetterlinge, Augsburg, 3: 37. TS: Papilio io Linnaeus, 1758.

 

io Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 472. TL: Sweden. Had been originally described as a good species.

·          geischa Stichel in Seitz, [1908] (Vanessa). Die Gross-Schmetterlinge der Erde, Stuttgart, 1: 201. TL: Japan. Had been originally described as a variety of io. D: Amur region, Ussuri region, Sakhalin.

·          io Linnaeus, 1758 (Papilio). D: European part; Ural; Caucasus; Transcaucasus; South Siberia; North Tian Shan; Dzhungaria.

 

Lepidoptera Indica, Madras, 4: 91. TS: Papilio canace Linnaeus, 1763.

 

canace Linnaeus, 1763 (Papilio). Amoenula academiae, 6: 406. TL: E China. Had been originally described as a good species.

·          charonides Stichel in Seitz, [1908] (Vanessa). Die Gross-Schmetterlinge der Erde, Stuttgart, 1: 206. TL: “Amur und Ussuri-Gebiet”. Had been originally described as a variety of canace. D: eastern part of Amur region, Ussuri region.

 

K. Svenska Vetensk. Acad. Handl., 1: 56. TS: Papilio urticae Linnaeus, 1758.

 

caschmirensis Kollar, 1848 (Vanessa). Butterflies of Kashmir, 4: 442; pl. 11, fig. 3, 4. TL: “in Ind. sup. Massuri, Caschmir”. Had been originally described as a good species.

·          nixa Grum-Grshimaïlo, 1890 (Vanessa). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 426. TL: “Hissar Mt.”, “Ходжа-Махметъ-Мазар” [Khodja-Makhmet-Mazar] – by the lectotype designation (Tuzov et al., 2000: 30). Had been originally described as a variety of urticae. D: Gissar; Darvaz; Alai; West Pamir.

 

connexa Butler & Fenton, 1881 (Vanessa). Proceedings of the zoological Society of London, 1881: 851. TL: Japan, Hokkaido: “Toshima”. Had been originally described as a variety of urticae. D: Sakhalin; South Kuriles.

·          connexa Butler & Fenton, 1881 (Vanessa).D: Sakhalin; South Kuriles.

 

ladakensis Moore, 1878 (Vanessa). Annales of the Magazin of Natural History, 5 (1): 227 - 228. TL: “Gorga, Changchenmo”. Had been originally described as a good species.

·          ladakensis Moore, 1878 (Vanessa). D: West Pamir.

 

rizana Moore, 1872 (Vanessa). Proceedings of the zoological Society of London, 1872: 559. TL: “Cheeni (9000 feet), Middle Kunawur, N.W.Himalaya”. Had been originally described as a good species.

·          rizana Moore, 1872 (Vanessa). D: Alai.

 

urticae Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 475. TL: Sweden. Had been originally described as a good species.

·          polaris Staudinger, 1871 (Vanessa). Catalog der Lepidopteren des europaeischen Faunengebietes, Berlin (Aufl. 2): 16. TL: Lappland. Had been originally described as a variety of urtica[e]. D: Northern part of Euroasia; Kamchatka; Magadan region; Yakutia.

·          urticae Linnaeus, 1758 (Papilio). D: All remaining territory of ex-USSR.

 

Subfamily Araschniinae Butler, 1869

Verzeichniss bekannter Schmetterlinge, Augsburg, 3: 37. TS: Papilio levana Linnaeus, 1758.

 

burejana Bremer, 1861. Bulletin de l’Academie des Sciences, Sankt-Petersburg, 3: 546. TL: “Bureja-Gebirge”. Had been originally described as a good species.

·          burejana Bremer, 1861. D: Amur region; Ussuri region; Sakhalin; South Kuriles.

 

levana Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 480. TL: Sweden. Had been originally described as a good species.

·          levana Linnaeus, 1758 (Papilio). D: European part; Ural; Caucasus; Siberia; Kamchatka; Amur region; Ussuri region; Sakhalin.

 

Subfamily Melitaeinae Tutt, 1896

 

Systematic Review of American Butterflies: 27. TS: Papilio phaeton Drury, [1773]

 

asiatica Staudinger, 1881 (Melitaea). Stettiner entomologische Zeitschrift, 42: 287. TL: Dzhungarian Alatau. Had been originally described as a variety of aurinia.

·          alexandrina Staudinger, 1887 (Melitaea). Stettiner entomologische Zeitschrift, 48: 52. TL: "Alexander-Gebirge". Had been originally described as a variety of aurinia. D: North Tian Shan.

·          asiatica Staudinger, 1881 (Melitaea). D: Dzhungaria.

·          narina Oberthür, 1909 (Melitaea). Études Lépidopterologie Compareé, 3: 230. TL: Naryn. Had been originally described as a variety of aurinia. D: Inner Tian Shan.

 

aurinia Rottemburg, 1775 (Papilio). Naturforscher, Halle, 6: 5. TL: Paris. Had been originally described as a good species.

·          aurinia Rottemburg, 1775 (Papilio). D: European part; Kazakhstan.

·          sibirica Staudinger, 1861 (Melitaea). Catalog der Lepidopteren des europäischen Faunengebietes, Berlin (Aufl. 1): 7. [nom. nov. pro Melitaea desfontainei Eversmann, 1851, non Melitaea desfontainei Boisduval, 1832]. TL: "Siberia". Had been originally established as a variety of artemis. D: Siberia.

 

banghaasi Stichel in Seitz, [1908] (Melitaea). Die Groß- Schmetterlinge der Erde. Die Groß- Schmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart, 1: 214. TL: "Kentei-Gebirge". Had been originally described as a subspecies of aurinia.

·          banghaasi Stichel in Seitz, [1908] (Melitaea). D: Altai.

 

davidi Oberthür, 1881 (Melitaea). Études d'Entomologie, 6: 52. TL: “Nord de la China”. Had been originally described as a variety of artemis.

·          davidi Oberthür, 1881 (Melitaea). D: Amur region.

·          tjutjujensis Higgins, 1950 (Euphydryas). Transactions of the Royak entomological Society of London, 101: 466. TL: "USSURI: Tjutjuje, Sichota Alin district; Transitional also from Anutschino, Chabarowsk, Sutschanski-Rudnik, Vladivostok". Had been originally described as a modification of siberica. D: Ussuri region.

 

iduna Dalman, 1816 (Melitaea). K. Svenska Vetensk. Acad. Handl., 37: 75. TL: Sweden. Had been originally described as a good species.

·          alferakyi Korshunov, 1996 (Euphydryas). Dopolnenija i ispravlenija k knige Dnevnye babochki asiatskoj chasti Rossii, Novosibirsk: 25. TL: "Якутия, 18 кмВСВп. Хандыга, верх. р. Хандыга" [Yakutia, 18 km ENE Khandyga, upper course of Khandyga river]. Had been originally described as a subspecies of iduna. D: Yakutia.

·          gorodinskii Churkin & Kolesnichenko, 2003 (Euphydryas). Helios, Moscow, 4: 107 – 109; pls. 6, 7, figs. 6, 12. TL: “Kamchatka, Bystrinskyi distr., Esso v., Uksichan R., 1200 m”. Had been originally described as a subspecies of iduna. D: TL.

·          iduna Dalman, 1816 (Melitaea). D: Polar part of European part and Siberia.

·          inexpectata Sheljuzhko, 1934 (Melitaea). Zeitschrift der Österreichischen entomologischen Vereins, 19: 1, figs. 1, 2. TL: Caucasus, Teberda, 2700 m. D: Caucasus.

·          sajana Higgins, 1950 (Euphydryas). Transactions of the Royal entomological Society of London, 101: 445. TL: "Sajan Mountains – Tunkinsk Weissberge", "south-eastern Altai Mountains". Had been originally described as a subspecies of iduna. D: Altai; Sayan.

·          semenovi Korshunov & Ivonin in Korshunov, 1996 (Euphydryas). Dopolnenija i ispravlenija k knige Dnevnye babochki asiatskoj chasti Rossii, Novosibirsk: 24. TL: "Хакассия, районг. Бобровой, ручей – притокр. Сарала" [Khakassia, area of Bobrovaya Mt. a brook near Sarala river]. Had been originally described as a subspecies of iduna. D: Kusnezky Alatau Mts.

 

intermedia Ménétriès in Schrenck, 1859 (Melitaea). Reisen und Forschungen im Amur-Lande, 2 (1): 22. TL: Amur. Had been originally described as a variety of maturna.

·          altaica Bang-Haas in Seitz, [1908] (Melitaea). Die Groß- Schmetterlinge der Erde. Die Groß- Schmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart, 1: 212. TL: “Altai”. D: Altai.

·          intermedia Ménétriès in Schrenck, 1859 (Melitaea). D: Amur region; Ussuri region.

·          konumensis Matsumura, 1927 (Melitaea). Insecta Matsumurasca, Sapporo, 2: 117; pl. 3, fig. 50. TL: Sakhalin: “Konuma”. Had been originally described as a subspecies of maturna. D: Sakhalin.

·          mongolica Staudinger, 1892 (Melitaea). Deutsche entomologische Zeitschrift Iris, 5: 321. TL: “Kentei”. Had been originally described as a variety of maturna. D: Tuva; Sayan; Praebaikal; Transbaikal.

 

maturna Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 480. TL: Europa. Had been originally described as a good species.

·          maturna Linnaeus, 1758 (Papilio). D: European part; West Siberia.

·          staudingeri Wnukowsky, 1929 (Melitaea). Zoologische Anzeiger, 83: 221 [nom. nov. pro Melitaea maturna uralensis Staudinger, 1871 nec Melitaea arduinna var. uralensis Eversmann, 1844]. TL: Ural. Had been originally described as a subspecies of maturna. D: South Ural; Altai; Dzhungaria.

 

merope de Prunner, 1798 (Papilio). Lepidoptera Pedemontana : 73. TL : Piedmont. Had been originally described as a good species.

·          altivolans Tuzov, 2000 (Euphydryas). Helios, Moscow, 1: 106. TL: “Altai, Severochuisky Mts., Sunkor Mt., 2500 m”. Had been originally described as a subspecies of merope. D: South Altai.

 

orientalis Herrich-Schäffer, 1845 (Melitaea). Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu Jakob Hübner's Sammlung europäischer Schmetterlinge. Umrisstafeln mit Erklärung. Nachträge. Systema Lepidopterorum. Index alphabetico-synonymicus ad Vol. I – V. Index universalis specierum et generum, 6: 1; Fign. 265, 266. TL: Amasia.

·          orientalis Herrich-Schäffer, 1845 (Melitaea). D: Little Caucasus; Transcaucasus.

·          sareptana Staudinger, 1871 (Melitaea). Catalog der Lepidopteren des europäischen Faunengebietes, Berlin (Aufl. 2): 17. TL: Sarepta. Had been originally described as a variety of aurinia. D: Caucasus; South of European part.

 

pellucida Christoph, 1893 (Melitaea). Deutsche entomologische Zeitschrift Iris, 6: 86. TL: East Caucasus, Daghestan, Kurush. Had been originally described as a variety of aurinia.

·          pellucida Christoph, 1893 (Melitaea). D: Caucasus.

 

provincialis Boisduval, 1828 (Melitaea). Index Methodicus, Paris: 17. TL: France, Provence. Had  been originally described as a variety of aurinia.

·          provincialis Boisduval, 1828 (Melitaea). D: Southern Azerbajdzhan.

 

Magazin für Insektenkunde, 6: 284. TS: Papilio cinxia Linnaeus, 1758.

 

acraeina Staudinger, 1886 (Melitaea). Stettiner entomologische Zeitschrift, 47: 233. TL: "Kokand" – by the lectotype (Kreuzberg, 1998: 38). Had been originally described as a good species.

·          acraeina Staudinger, 1886 (Melitaea). D: Fergana valley.

 

ala Staudinger, 1881 (Melitaea). Stettiner entomologische Zeitschrift, 42: 288. TL: "Lepsa". Had been originally described as a variety of didyma.

·          ala Staudinger, 1881 (Melitaea). D: Tian Shan.

·          irtyschica Lukhtanov, 1999 (Melitaea). Atalanta, Würzburg, 30 (1/4): 144 – 145. TL: “Kasachstan, Ust-Kamenogorsk-Gebiet, Ulbinski-Gebirge, Oktjabrsk, 420 m.” Had been originally described as a subspecies of ala. D: southern part of Ulbinsky mountain ridge, Irtysh valley and Bukhtarma valley in West Altai.

·          zaisana Lukhtanov, 1999 (Melitaea). Atalanta, Würzburg, 30 (1/4): 143 – 144. TL: “Kasachstan, Ust-Kamenogorsk-Gebiet, Süd-Altai, Kurtschum-Gebirge, zwischen Kurtschum und Kalguty, 600 m.” Had been originally described as a subspecies of ala. D: southern parts of Narymsky, Kurchumsky and Asutau mountains, Bukombai Mts. in South Altai.

 

alrashid Higgins, 1941 (Melitaea). Transactions of the Royal entomological Society of London, 91: 259 – 260. TL: “Western part of the Gissar Mountain System, close to Samarcand” (Churkin, 2000: 92). Had been originally described as a subspecies of amrisia.

·            alrashid Higgins, 1941 (Melitaea). D: East Gissar.

·            garun Churkin, 2000 (Melitaea). Helios, Moscow, 1: 94; fig. 3 f. TL: “Kirgizstan, Matcha, eastern edge of the Turkestansky Mts., upper stream of Ak-Terek River, Noo-Dzhaiko, 3550 – 3700 m”. Had been originally described as a subspecies of alrashid. D: West Gissar.

 

ambrisia Higgins, 1935 (Melitaea). Entomologist, London, 68: 112. TL: "Marghilan". Had been originally described as a good species.

·          ambrisia Higgins, 1935 (Melitaea). D: Alai; Darvaz.

·          avicaena Churkin, 2000 (Melitaea). Helios, Moscow, 1: 91 – 92; figs. 1 b, 2 b, g, 3 d. TL: “Uzbekistan, Turkestansky Range, Kumbel Pass, Яффьштылн res., 3000 m”. Had been originally described as a subspecies of ambrisia. D: Gissar.

 

arcesia Bremer, 1864 (Melitaea). Bulletin  de la Academie impériale des Sciences, Saint-Pétersbourg, 6: 545. TL: “Norden des Baikal und in Daurien”. Had been originally described as a good species.

·          arcesia Bremer, 1864 (Melitaea). D: Transbaikal; Amur region; Ussuri region.

·          dabanica Barantchikov, 1979 (Melitaea). Fauna lesov bassejna ozera Baikal, Novosibirsk: 109 – 123. TL: "Хамар-Дабан, верховьяр. ЛеваяМишиха, субальпийскийпояс" [Khamar-Daban, upper course of Levaja Mishikha river, subalpine zone]. Had been originally described as a subspecies of arcesia. D: TL.

·          minor Elwes, 1899 (Melitaea). Transactions of the Royal entomological Society of London, 1899: 334. TL: Altai, Chuya Mts., 1950 – 2400 m. Had been originally described as a variety of arcesia. D: Altai; Sayan; Tuva.

 

arduinna Esper, [1784] (Papilio). Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen, Erlangen, 1 (2): 169; Tab. 87. TL: Kamyshin in the Volgograd region. Had been originally described as a good species.

·          arduinna Esper, [1784] (Papilio). D: South of European part; Caucasus; Kazakhstan; North Tian Shan; Alai; Darvaz; West Pamir.

·          evanescens Staudinger, 1886 (Melitaea). Stettiner entomologische Zeitschrift, 47: 231. TL: North Persia. Had been originally described as a variety of arduinna. D: Gissar; Kopet Dagh.

·          rhodopensis Freyer, 1836 (Papilio). Neue Beiträge zur Schmetterlingskunde mit Abbildungen nach der Natur, Augsburg, 3: 3; Taf. 193, Fig. 1. TL: "Türkei". Had been originally described as a good species. D: Transcaucasus.

 

asteroida Staudinger, 1881 (Melitaea). Stettiner entomologische Zeitschrift, 42: 292. TL: Dzhungarian Alatau. Had been originally described as a good species.

·          asteroida Staudinger, 1881 (Melitaea). D: Dzhungaria.

·          clara Staudinger, 1887 (Melitaea). Stettiner entomologische Zeitschrift, 48: 55. TL: Kouldja. Had been originally described as a subspecies of asteroida. D: Inner Tian Shan.

·          pallida Staudinger, 1901 (Melitaea). Catalog der Lepidopteren des palaearktischen Faunengebietes, Berlin (Aufl. 1): 33. TL: Osh. Had been originally described as a subspecies of asteroida. D: East Alai.

 

athene Staudinger, 1881 (Melitaea). Stettiner entomologische Zeitschrift, 42: 266. TL: Saisan. Had been originally described as a good species.

·          athene Staudinger, 1881 (Melitaea). D: Dzhungaria; SW Altai; North Nian Shan: mountain ridges Kungey Ala Too, Terskey Ala Too, Ketmen, Kirghiz (east).

 

avinovi Sheljuzhko, 1914 (Melitaea). Deutsche entomologische Zeitschrift Iris, 26: 20. TL: "Ischkaschim". Had been originally described as a subspecies of arduinna.

·          avinovi Sheljuzhko, 1914 (Melitaea). D: Only known from the Ishkashimsky mountain ridge.

 

cassandra Kolesnichenko & Churkin, 2001 (Melitaea). Helios, Moscow, 2: 143 – 149; figs. 1 – 3, pl. 14, figs. 1 – 4. TL: “Kirgizstan, Suusamyr Mts., Kokemeren r., 5 km N Kyzyl-Oi, 1800 – 2000 m”. Had been originally described as a good species.

·            cassandra Kolesnichenko & Churkin, 2001 (Melitaea). D: Suusamyr valley, mountains around Suusamyr valley, southern slopes of Kungey Ala-Too Mts., Orto-Tokoi Lake vicinity.

 

catapelia Staudinger, 1886 (Melitaea). Stettiner entomologische Zeitschrift, 47: 231. TL: "Umgegend von Samarkand". Had been originally described as a variety of trivia.

·          bactriana Ju.Ju.Stshetkin, 1984 (Melitaea). Zoological Journal, Moscow, 63: 1823. TL: "окрестностиг. Калининабад (ТаджикскаяССР), угребняхребтаТабакчи,… 1200 м" [Kalininabad vicinity (Tadzhik SSR), Tabakchi Mts., 1200 m]. Had been originally described as a subspecies of catapelia. D: South Gissar (ridges of the South-Tadzhikistanian tectonic depression).

·          catapelia Staudinger, 1886 (Melitaea). D: West Gissar.

·          chorasana Ju.Ju.Stshetkin, 1984 (Melitaea). Zoological Journal, Moscow, 63: 1824. TL: "северо-восточныйИран, окрестностиг. Мешхеда, …830 м" [North-East Iran, Meshkhed vicinity, 830 m]. Had been originally described as a subspecies of catapelia. D: Kopet Dagh.

·          nativa Tuzov, 2000 (Melitaea). A guide to the butterflies of Russia and adjacent territories. Sofia, 2: pl. 40, figs. 31 – 33. TL: “Tadjikistan, W. Pamirs, Vanchsky Mts., Lyangar, 2800 m”. Had been originally described as a subspecies of trivia. D: West Pamir: Vanchsky Mts.

·          petri Ju.Ju.Stshetkin, 1984 (Melitaea). Zoological Journal, Moscow, 63: 1824 – 1825. TL: "северныйсклонцентральнойчастихребтаПетраПервого, среднеетечениер. Дараи-Назарак, в 14 кмюго-западнееТаджикабада, …2400 м" [northern slope of the central part of Peter the Great mountain ridge, middle course of Darai-Nazarak river, 14 km SW Tadzhikabad, 2400 m]. Had been originally described as a subspecies of catapelia. D: East Gissar; Darvaz.

·          nativa Tuzov in Tuzov et al., 2000 (Melitaea) nom. nov. pro shugnana Stshetkin, 1984, nec shugnana Sheljuzhko, 1928. A guide to the butterflies of Russia and adjacent territories, Sofia, 2: 62. TL: "западнаячастьШугнанскогохребта, близг. Хорог, ущельеСангоудара, …3020 м" [Western part of Shugnan Mts., near Khorog, Sangoudara valley, 3020 m]. Had been originally described as a subspecies of catapelia. D: West Pamir: Ishkashimsky and Shugnansky Mts.

 

chitralensis Moore, 1901 (Melitaea). Lepidoptera Indica, Madras, 5: 9. TL: "Chitral". Had been originally described as a good species.

·          shugnana Sheljuzhko, 1929 (Melitaea). Mitteilungen der Münchener entomologischen Gesellschaft, 19: 355. TL: "Pamir, Chorog". Had been originally described as a subspecies of didyma. D: West Pamir.

 

cinxia Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 480. TL: Sweden. Had been originally described as a good species.

·          amardea Grum-Grshimailo, 1895 (Melitaea). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 29: 291. TL: "in regione alpina montium Elburs (Demavend)". Had been originally described as a variety of cinxia. D: Kopet Dagh.

·          aversa Churkin & Kolesnichenko, 2002 (Melitaea). Helios, Moscow, 3: 137 – 141; figs. 1 a, b, e, f, 2 a – d; pl. 11, figs. 1 – 6. TL: “Kirgizstan, Tian-Shan, Suusamyr Mts., 35 km SE Alabel Pass, Aram-Suu River, 2500 – 2700 m”. Had been originally described as a subspecies of cinxia. D: Suusamyr Mts., Baidulu Mts.

·          cinxia Linnaeus, 1758 (Papilio). D: European part; Caucasus; Crimea.

·          clarissa Staudinger, 1901 (Melitaea). Catalog der Lepidopteren des palaearktischen Faunengebietes, Berlin (Aufl. 1): 29. TL: Asia Minor. Had been originally described as a variety of cinxia. D: Transcaucasus.

·          oasis Huang & Murayama, 1992 (Melitaea). Tyô to Ga, Tokyo, 43: 7; fig. 20. TL: "Jiadengyu, Altai, 1380 – 1600 m, grassland in forest zone". Had been originally described as a subspecies of cinxia. D: SW Altai.

·          tschujaca Seitz, 1909 (Melitaea). Die Groß- Schmetterlinge der Erde. Die Groß-Schmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart, 1: 215. TL: "Altai". Had been originally described as a subspecies of cinxia. D: South Siberian mountains; Dzhungaria; Saur; Tarbagatai.

 

diamina Lang, 1789 (Papilio). Verzeichniss der Schmetterlinge von Gegend des Augsburgs (ed. 2): 44. TL: "Augsburg". Had been originally described as a good species.

·          badukensis Alberty, 1969 (Melitaea). Deutsche entomologische Zeitschrift, 16: 193; Taf. 1, Abbn. 5, 6. TL: Baduk sea in Caucasus. Had been originally described as a subspecies of dictynna. D: Caucasus.

·          erycina Lederer, 1853 (Melitaea). Verhandlungen des zoologisch-botanischen Vereins, Wien, 3: 357. TL: “Ustbuchtarminsk”. Had been originally described as a variety of dictynna Esper, [1777]. D: Siberia; Far East.

·          hebe Borkhausen, 1793 (Papilio). Rheinisches Magazin, 1: 270. TL: Europa. Had been originally described as a good species. D: European part; North Kazakhstan.

 

didyma Esper, [1779] (Papilio). Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen, Erlangen, 1: 365. TL: Germany. Had been originally described as a good species.

·          ambra Higgins, 1941 (Melitaea). Transactions of the Royal entomological Society of London, 91: 208 – 209 [nom. nov. pro Melitaea didyma dschungarica Oberthür, 1909, nec Melitaea sibina dschungarica Grum-Grshimailo, 1895]. TL: "Dzungaria". Had been originally described as a subspecies of didyma. D: Dzhungaria.

·          altaica Grum-Grshimailo, 1893 (Melitaea). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 27: 127. TL: "Alati". Had been originally described as a variety of didyma. D: Altai; Tuva; Saur; Tarbagatai.

·          didyma Esper, [1779] (Papilio). D: Middle  of European part.

·          manoni Hanus, 1996 (Melitaea). Atalanta, Würzburg, 27: 207; pl. 4, figs. 2 [non 4!], 4 [non 5!], 6 [all "non!" – by the author's remarks on the reprint of original description which I received from J.Hanus]. TL: "Kirghizia, Kadzhi Say, 1610 m on the south shore of Issyk-kul". Had been originally described as a subspecies of didyma. D: Terskey Ala Too Mts.

·          neera Fischer de Waldheim, 1840 (Melitaea). Bulletin de la Societe impériale des Naturalistes de Moscou, 13 (3): 81. TL: Volga. Had been originally described as a variety of didyma. D:  South of European part.

·          pseudolatonigena Yakovlev, 2002 (Melitaea). Helios, Moscow, 3: 102 – 105; pl. 7, figs. 1 – 12. TL: “N Tuva, the Uyukskii Mt. Range, 30 km N of Kyzyl, 800 m”. Had been originally described as a subspecies of didyma. D: TL.

·          turkestanica Sheljuzhko, 1929 (Melitaea). Mitteilungen der Münchener entomologischen Gesellschaft, 19: 354. TL: "Margelan, Osch, Usgent". Had been originally described as a variety of didyma. D: Middle Asiatic mountains.

 

didymoides Eversmann, 1847 (Melitaea). Bulletin de la Société impériale des Naturalistes de Moscou, 20 (2): 67; pl. 1, figs. 3, 4. TL: "Kiachtam". Had been originally described as a good species.

·          didymoides Eversmann, 1847 (Melitaea). D: Tuva (Malyj Enisej river); Sayan (Mondy); South Praebaikal; South and East Transbaikal; Zea river valley; South Ussuri region.

 

elisabethae Avinov, 1910 (Melitaea). Horae Societatis entomologicae Rossicae, Sankt-Petersburg, 39: 235; Tab. 14; fig. 6 et 7. TL: "в горахъ южнhе Аличура" [in the mountains souther of Alichur]. Had been originally described as a good species.

·          elisabethae Avinov, 1910 (Melitaea). D: West Pamir.

 

543. enarea Fruhstorfer, 1916 (Melitaea). Archiv Naturgeschichten, 82 A (2): 11. TL: "Garm, Pamir (Peter the Great Mts.)". Had been originally described as a subspecies of didyma.

·          enarea Fruhstorfer, 1916 (Melitaea). D: Gissar, Darvaz.

·          gromenkoi Kolesnichenko, 2000 (Melitaea). A guide to the butterflies of Russia and adjacent territories, Sofia, 2: pl. 38, figs. 29, 30. TL: “Kirghizia, Alaisky Mts. (east), Kyzyldzhar, 2,300 m a.s.l.” Had been originally described as a subspecies of enarea. D: East Alai.

 

enoch Higgins, 1941 (Melitaea). Transactions of the Royal entomological Society of London, 91: 337; pl. 14, fig. 4. TL: "Askabad". Had been originally described as a modification of phoebe.

·          enoch Higgins, 1941 (Melitaea). D: Kopet Dagh; South Transcaucasus.

 

fergana Staudinger, 1882 (Melitaea). Berliner entomologische Zeitschrift, 26: 168. TL: [Alai] – by the lectotype designation (Higgins, 1941: 256; pl. 7, fig. 3). Had been originally described as a good species.

·          darvasica Avinoff in Higgins, 1941 (Melitaea). Transactions of the Royal entomological Society of London, 91: 257. TL: "Бухара, Дарваз, пер. Висхарви" [Bukhara, Darvaz, Viskharvi Pass] – by the lectotype (Churkin & Tuzov, 2000b: 41). Had been originally described as a subspecies of fergana. D: Alai; Darvaz.

·          fergana Staudinger, 1882 (Melitaea). D: Inner Tian Shan; Alai.

·          grumi Churkin & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 44 – 45; figs. 2 e, 3 h, 7 a. TL: “Tadjikistan, East Pamirs, Mynkhadzhir Mts., … 4100 – 4300 m”. Had been originally described as a subspecies of fergana. D: East Pamir: Mynkhadzhir Mts.

·          jacobsoni Avinoff in Higgins, 1941 (Melitaea). Transactions of the Royal entomological Society of London, 91: 257; pl. 7, figs. 2, 8. TL: "Pamirs: Maz River". Had been originally described as a subspecies of fergana. D: East Pamir.

·          ketmeana Lukhtanov, 1999 (Melitaea). Atalanta, Würzburg, 30 (1/4): 145 – 146. TL: “Kasachstan, Alma-Ata-Gebiet, Ketmen-Gebirge, Kolshat”. Had been originally described as a subspecies of fergana. D: Ketmen Mts., Kungey Ala-Too Mts.

·          khantengri Churkin & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 50 – 51; figs. 2 h, 5 f, 7 d. TL: “Kirghizia, Sary-Dzas R., Kaindy-Katta Mts., Tashkoro loc., 3100 m”. Had been originally described as a subspecies of fergana. D: only known from the type locality.

·          maracandica Staudinger, 1882 (Melitaea). Berliner entomologische Zeitschrift, 26: 169. TL: "Hazret Sultan Gebirge". Had been originally described as a variety of fergana. D: West Pamir; Darvaz; Gissar.

·          oxuana Lukhtanov, 1999 (Melitaea). Atalanta, Würzburg, 30 (1/4): 147. TL: “Tadschikistan, Ost-Pamir, Sarykol-Gebirge, Dunkeidyk [sic! right: Dunkeldyk], 3750 m.” Had been originally described as a subspecies of fergana. D: East Pamir.

·          paradoxa Churkin & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 48 – 50; figs. 1 c, 2 f, g, 4 c, d. TL: “Kazakhstan, 170 km W. Alma-Ata, Dzhetyzhol Mts., 1400 m”. Had been originally described as a subspecies of fergana. D: Zhetizhol Mts. in North Tian Shan.

·          ssp? D: Only known from Kirghiz mountain ridge, vicinity of Bishkek, Chon Kuurchak valley.

·          splendida Churkin & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 39 – 40; figs. 2 a, 3 b, 6 a. TL: “Uzbekistan, Turkestansky Mts., Zaaminsky Reservation near near Kumbel Pass, 3200 m”. Had been originally described as a subspecies of fergana. D: Gissar.

·          terskeana Lukhtanov, 1999 (Melitaea). Atalanta, Würzburg, 30 (1/4): 146. TL: “Kirgisien, Terskey-Alatoo, Barskaun-Pass, 3400 m.” Had been originally described as a subspecies of fergana. D: Terskey Ala-Too Mts.

·          vladislavi Churkin & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 45 – 48; figs. 3 f, g, 4 a, 5 e, 6 c, d. TL: “Kirghizia, Inner Tian-Shan, At-Bashi Mts., 5 km У Bosogo v., 2200 – 2500 m”. Had been originally described as a subspecies of fergana. D: Inner Tian Shan.

 

hafiz Higgins, 1941 (Melitaea). Transactions of the Royal entomological Society of London, 91: 225; pl. 4, fig. 5. TL: "In the north of Persia". Had been originally described as a forma of persea.

·          hafiz Higgins, 1941 (Melitaea). D: South Transcaucasus: Talysh.

 

infernalis Grum-Grshimailo, 1891 (Melitaea). Horae Societatis entomologicae Rossicae, Sankt-Petersburg, 25: 455. TL: "Tian-Schan / Boro Khoro", “Унканъ” [Unkan] – by the lectotype (Churkin & Kolesnichenko, 2002: 145). Had been originally described as a variety of saxatilis Christoph, 1873.

·          carmen Churkin & Kolesnichenko, 2002 (Melitaea). Helios, Moscow, 3: 148 – 150; figs. 1 c, d, 2 c, d, 3 c, d, 4 b, pl. 10, figs. 5 – 8. TL: “Kasakhstan, Dzhungaria, Burchansarytau Mts., 15 km W Enbekshi, 2700 m”. Had been originally described as a subspecies of infernalis. D: SW Dzhungaria.

 

interrupta Skala, 1907 (Melitaea). Entomologische Zeitschrift, Guben, 20: 311. TL: Nikolsburg. Had been originally described as an aberration of didyma.

·       interrupta Skala, 1907 (Melitaea). D: Transcaucasus; Caucasus; Praecaucasian part of Russia (Stavropolsky Kray).

·       kendevana Gross & Ebert, 1975 (Melitaes). Journal of the Entomological Society of Iran, Teheran, Suppl. 1: 37. TL: Demavend, Iran. Had been originally described as a subspecies of didyma. D: Kopeth-Dagh.

 

kotshubeji Sheljuzhko, 1929 (Melitaea). Mitteilungen der Münchener entomologischen Gesellschaft, 19: 364. TL: "Jugum Petri Magni (Buchara or.): Tuptshek". Had been originally described as a subspecies of ala. D: Darvaz; Alai; northern ridges of West Pamir.

·       kotshubeji Sheljuzhko, 1929 (Melitaea). D: Darvaz.

·       bundeli Kolesnichenko, 2000 (Melitaea). A guide to the butterflies of Russia and adjacent territories. Sofia, 2: pl. 38, figs. 31, 32. TL: “Kirghizia, Alaisky Mts. (east), Kyzyldzhar, 2500 m a.s.l.” Had been originally described as a subspecies of kotshubeji. D: Alai; northern ridges of Pamir.

 

latonigena Eversmann, 1847 (Melitaea). Bulletin de la Société impériale des Naturalistes de Moscou, 20 (3): 66; pl. 1, fig. 1, 2. TL: "in provincia Irkutskensi". Had been originally described as a good species.

·          atrata Higgins, 1935 (Melitaea). Entomologist, London, 68: 112. TL: "Sajan Mountains". Had been originally described as a subspecies of latonigena. D: Altai (without Ukok plateau); Sajan; Tuva.

·          latonigena Eversmann, 1847 (Melitaea). D: Praebaikal; Transbaikal.

·          polaris Grum-Grshimailo, 1899 (Melitaea). Annales Misei Zoologici, Sankt-Petersburg, 4: 459. TL: "Verchojansk".Had been originally described as a variety of didyma. D: Far East.

·          ukoka Korshunov, 1998 (Melitaea). Novye opisanija i utochnenia dlya knigi Dnevnye babochki asiatskoj chasti Rossii, Novosibirsk: 19. TL: “наплоскогорьеУкоквбассейнер. Калгуты” [on the Ukok plateau in the Kalguty river basin]. Had been originally described as a subspecies of latonigena. D: Ukok plateau in SW Altai.

 

ludmilla Churkin, Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 74 – 76; figs. 1 a, 8 a. TL: “Tian-Shan, Kirghizsky Range, Uzungyr Mts., Alamedin R., 3000 - 3200 m”. Had been originally described as a good species.

·            ludmilla Churkin, Kolesnichenko & Tuzov, 2000 (Melitaea). D: North Tian-Shan: Kirghiz Mts.

·            serena Churkin, Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 76 – 77; figs. 1 b, 5 b, h, j6 b, 7 c. TL: “Tian-Shan, Talassky Alatau, Otmek Pass, 3400 m”. Had been originally described as a subspecies of ludmilla. D: West Tian Shan: Talassky Mts.

 

lunulata Staudinger, 1901 (Melitaea). Catalog der Lepidopteren des palaearktischen Faunengebietes (Aufl. 1), Berlin: 30. TL: “Taschkend” (Lukhtanov, 1999: 148). Had been originally described as a variety of saxatilis.

·          merke Lukhtanov, 1999 (Melitaea). Atalanta, Würzburg, 30 (1/4): 147 – 148. TL: “Kasachstan, Kirgisisches Gebirge, Merke Fl., 2900 m.” Had been originally described as a subspecies of lunulata. D: Kirghiz Mts.

·          lunulata Staudinger, 1901 (Melitaea). D: West Tian Shan; Kirghiz Mts.

 

lutko Evans, 1932 (Melitaea). Identification of Indian butterflies, Madras (ed. 2): 185. TL: "Chitral". Had been originally described as a subspecies of robertsi.

·          mimetica Higgins, 1940 (Melitaea). Entomologist, London, 73: 52. TL: "Baluchistan: Khojak, …6500 ft". Had been originally described as a subspecies of lukto [sic !]. D: Badkhyz in South Turkmenia (Devyatkin & Tuzov, 1988).

 

minerva Staudinger, 1881 (Melitaea). Stettiner entomologische Zeitschrift, 42: 289. TL: Dzhungarian Alatau. Had been originally described as a good species.

·          acerba Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 118; fig. 4. TL: “Mts. Tschatkalensis merid., lacus Sary-Tschilek, 2400 m”. Had been originally described as a subspecies of minerva. D: West Tian Shan: Chatkalsky Mts.

·          balba Evans, 1912 (Melitaea). Journal of the Bombay Natural history Society, 21: 582. TL: "Chitral". Had been originally described as a race of balbita. D: East Pamir.

·          balbina Tytler, 1926 (Melitaea). Journal of the Bombay natural History Society, 31 (3): 583. TL: “Shandur Pass between Gilgit and Chitral at 12,200’”. Had been originally described as a subspecies of balba. D: Muzkol, Pshartsky and the eastern part of Yuzhno-Alichursky mountain ridges.

·          minerva Staudinger, 1881 (Melitaea). D: North Tian Shan.

·          palamedes Grum-Grshimaïlo, 1890 (Melitaea). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 4: 436. TL: “Гульча” [Gultcha] – by the lectotype (Kolesnichenko & Churkin, 2003: 306). Had been originally described as a variety of minerva. D: Alai; Darvaz; Gissar; West Pamir.

·          pseudotersa Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 114 – 115; fig. 1. TL: “Baidullu Mts., 35 km S Dolon Pass, Kara-Unkur River, 1800 m””. Had been originally described as a subspecies of minerva. D: Inner Tian Shan.

·          repens Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 115 – 116; fig. 2. TL: “Kara-Tau Mts., near Kentau, 800 m”. Had been originally described as a subspecies of minerva. D: West Tian Shan: Syrdarjinski Karatau Mts.

·          tersa Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 116 – 117; fig. 3. TL: “Uzbekistan, 70 km from Tashkent, Karzhantau Mts., Aktash v., 1900 m”. Had been originally described as a subspecies of minerva. D: West Tian Shan: Karzhantau Mts.

 

nadezhdae Sheljuzhko, 1912 (Melitaea), stat.n. Deutsche entomologische Zeitschrift Iris, 26: 137. TL: "Pamirskij Post". Had been originally described as a subspecies of didyma.

Remark. Agrees practically to all modern works on the butterflies of Central Asia, the taxon nadezhdae it is considered as a subspecies of  mixta. Between that the name nadezhdae Sheljuzhko is published on one half-year earlier than mixta, therefore here, were guided a principle of a priority, I fix for nadezhdae the specific status and I translate mixta as a subspecies of nadezhdae: Melitaea nadezhdae mixta Evans, 1912, stat.n.

·          nadezhdae Sheljuzhko, 1912 (Melitaea). D: East Pamir.

·          problematica Sheljuzhko, 1929 (Melitaea). Mitteilungen der Münchener entomologischen Gesellschaft, 19: 357. TL: West Pamir. Had been originally described as a subspecies of didyma. D: West Pamir; Darvaz.

 

ninae Sheljuzhko, 1935 (Melitaea). Mitteilungen der Münchener entomologischen Gesellschaft, 25: 27. TL: Chimgan, West Tian Shan. Had been originally described as a subspecies of ala.

Remark. The name pseudoala, used by modern entomologists, can not be used in quality of specific or subspecific, as was initially applied as infrasubspecific: Melitaea didyma turkestanica ab. ♀ pseudoala Sheljuzhko, 1928. Later L.Sheljuzhko has described the subspecies Melitaea ala ninae Sheljuzhko, 1935. This name is satisfied to all requirements of suitability, therefore only with it, as the oldest suitable name for the taxon, designated earlier as pseudoala, can be used in the biological nomenclature. The name rosea Higgins, 1938, offered by two years after the name of L.Sheljuzhko, is a younger subjective synonym of the name ninae.

·          ninae Sheljuzhko, 1935 (Melitaea). D: West and North Tian Shan.

 

pallas Staudinger, 1886 (Melitaea). Stettiner entomologische Zeitschrift, 42: 235. TL: "südlichen Alai". Had been originally described as a variety of minerva.

·          interpres Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 121; fig. 7. TL: “Tian-Shan Mts. Centr., Naryn”. Had been originally described as a subspecies of pallas. D: Inner Tian Shan: Naryn vicinity.

·          iracunda Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 119; fig. 5. TL: “W. Tian-Shan, Talassky Alatau, Kara-Bura River, Kara-Bura Pass, 3000 m”. Had been originally described as a subspecies of pallas. D: West Tian Shan.

·          koitezek Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 124; fig. 10. TL: “Pamirs, Koi-Tezek Pass, 4400 m”. Had been originally described as a subspecies of pallas. D: East Pamir: South-Alichursky Mts.

·          mendax Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 120; fig. 6. TL: “Kirgizsky Mts., Alamedyn River (“Александровкий хребет, р. Аламедын”)”. Had been originally described as a subspecies of pallas. D: North Tian-Shan: Kirghiz Mts.

·          pallas Staudinger, 1886 (Melitaea). D: Alai; Beleuli Mts.

·          pseudobalbina Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 122 – 124; fig. 9. TL: “SE. Pamirs, Sarykol’sky Mts., Tokhtamysh, 4000 m”. Had been originally described as a subspecies of pallas. D: East Pamir: Sarykolsky Mts.

·          vashchenkoi Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 122. TL: “Inn. Tian-Shan Mts., At-Bashi Mts., 5 km E Bosogdo v., 2500 – 2800 m”. Had been originally described as a subspecies of pallas. D: Inner Tian Shan: At-Bashi Mts. and environs of the Lake Chatyrkul.

 

permuta Higgins, 1941 (Melitaea). Transactions of the Royal entomological Society of London, 91: 250. TL: "from the Hazret-Sultan Mountains", "Aulie-Ata". Had been originally described as a modification of enarea.

·            permuta Higgins, 1941 (Melitaea). D: Gissar.

 

persea Kollar, [1849] (Melitaea). Denkschriften der Akademie den Wissenschaften, Wien, 1: 52. TL: "Shiraz". Had been originally described as a good species.

·          araratica Verity, 1929 (Melitaea). Entomological Record, London, 41: 113. TL: "Mount. Ararat". Had been originally described as a subspecies of didyma. D: Transcaucasus.

·          persea Kollar, [1849] (Melitaea). D: Kopet Dagh; South West Tadzhikistan.

·          ssp? D: West Tian Shan; Gissar (Tuzov, 1993: 48).

 

phoebe [Denis & Schiffermüller], [1775] (Papilio). Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend, Wien: 179. TL: Vienna. Had been originally described as a good species.

·          aetherea Eversmann, 1851 (Melitaea). Entomographia imperii Rossici, Sankt-Petersburg, 5: 73; pl. 5, fig. 5, 6. TL: South Russia. Had been originally described as a subspecies of phoebe. D: South of European part; North Kazakhstan; Crimea; Caucasus.

·          changaica Seitz, 1909 (Melitaea). Die Groß- Schmetterlinge der Erde. Die Groß-Schmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart, 1: 217. TL: "Changai". Had been originally described as a subspecies of phoebe. D: Amur region.

·          ornata Christoph, 1893 (Melitaea). Deutsche entomologische Zeitschrift Iris, 6: 87. TL: South Ural. Had been originally described as a variety of phoebe. D: South Ural; West Siberia.

·          ottonis Fruhstorfer, 1917 (Melitaea). Archiv Naturgeschichten, 82 A (2): 2 [nom. nov. pro Melitaea phoebe var. caucasica Staudinger, 1870 nec Melitaea didyma var. caucasica Staudinger, 1861]. TL: "Armenia". Had been originally described as a variety of phoebe. D: Little Caucasus; Transcaucasus.

·          saturata Staudinger, 1892 (Melitaea). Deutsche entomologische Zeitschrift Iris, 5: 323. TL: Kentei. Had been originally described as a variety of phoebe. D: Dzhungaria; Saur; Tarbagatai;  Altai; Transili Alatau Mts.; Ketmen Mts.; Kungey Ala Too Mts.

·          tungana Seitz, 1909 (Melitaea). Die Groß-Schmetterlinge der Erde. Die Groß-Schmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart, 1: 216. TL: "Sajan". Had been originally described as a subspecies of phoebe. D: Sayan; Tuva; Praebaikal; Transbaikal.

·          tungusa Herz, 1898 (Melitaea). Deutsche entomologische Zeitschrift Iris, 11: 240. TL: "Witim", "Wilui". Had been originally described as a variety of phoebe. D: Middle Siberia; Middle Far East.

 

protomedia Ménétriès, 1858 (Melitaea). Bulletin de l’Academie des Sciences, Saint-Pétersbourg, 17: 214. TL: Lover Amur. Had been originally described as a good species.

·          protomedia Ménétriès, 1858 (Melitaea). D: Middle Amur region.

·          regama Fruhstorfer, 1915 (Melitaea). Societas Entomologica, 30: 58. TL: "Chang-Yang" in West China. Had been originally described as a subspecies of protomedia. D: Ussuri region.

 

punica Oberthur, 1876 (Melitaea).  Etudes d’Entomologie, 1: 25. TL: Lambessa, Algeria. Had been originally described as a variety of phoebe.

·            amanica Rebel, 1917 (Melitaea). Sitzb. k. Akad.-Wiss. Wien, 126: 252. TL: “Amanus”. Had been originally described as a subspecies of phoebe. D: Azerbajdzhan, Talysh.

·            ssp? D: Volgograd region on Southern Russia.

 

romanovi Grum-Grshimailo, 1891 (Melitaea). Horae Societatis entomologicae Rossicae, Sankt-Petersburg, 25: 454. TL: "Dscharchar" (Staudinger, 1892: 373). Had been originally described as a good species.

·          puella Higgins, 1941 (Melitaea). Transactions of the Royal entomological Society of London, 91: 274; pl. 9, fig. 11. TL: "Amdo: Mougjik, Hoang-Ho Valley". Had been originally described as a modification of romanovi. D: South Altai (Ukok); South Transbaikal (rivers Selenga, Onon, lakes Bolshoj Chindant, Borun-Torej, Zun-Torej).

 

scotosia Butler, 1878 (Melitaea). Cistula Entomologica, 2: 282. TL: "Tokyo". Had been originally described as a good species.

·          butleri Higgins, 1940 (Melitaea). Entomologist, London, 73: 53. TL: "Pekin". Had been originally described as a subspecies of scotosia. D: South and moddle part of Ussuri region.

 

shandura Evans, 1924 (Melitaea). Journal of the Bombay natural History, 30: 90. TL: "Shandur". Had been originally described as a good species.

·          pavlitzkajana Sheljuzhko, 1943 (Melitaea). TL: "Darvaz, Jugum Vantsh mts, Gushchon-Pass". Had been originally described as a subspecies of didyma. D: Pamir; Darvaz.

 

sibina Alphéraky, 1881 (Melitaea). Horae Societatis entomologicae Rossicae, Sankt-Petersburg, 16: 400. TL: "Kouldjà". Had been originally described as a variety of phoebe.

·          dshungarica Grum-Grshimailo, 1895 (Melitaea). Horae Societatis entomologicae Rossicae, Sankt-Petersburg, 29: 291. TL: "In Dshungaria ad Dshincho". Had been originally described as a variety of sibina. D: Dzhungaria.

·          rama Higgins, 1941 (Melitaea). Transactions of the Royal entomological Society of London, 91: 349; pl. 15, fig. 5. TL: "Alai". Had been originally described as a modification of sibina. D: All Middle Asiatic mountains.

·          sibina Alphéraky, 1881 (Melitaea). D: Ili region; Ketmen Mts; Saur; Tarbagatai.

·          weiwueria Huang & Murayama, 1992 (Melitaea). Tyô to Ga, Tokyo, 43: 7; fig. 19. TL: "Jiadengyu, Altai, 1400 m, grassland in forest zone". Had been originally described as a subspecies of scotosia. D: SW Altai.

 

solona Alphéraky, 1881 (Melitaea). Horae Societatis entomologicae Rossicae, Sankt-Petersburg, 16: 404; pl. 14, figs. 14, 15. TL: "Tian-Chian, Jouldousse". Had been originally described as a forma of minerva.

·          evadne Hemming, 1934 (Melitaea) (Melitaea asteroida evadne nom. nov. pro Melitaea pallida Staudinger, 1901, nec Melitaea pallida Edwards, 1864). Stylops, 3: 98. TL:  Osh. Had been originalle described as a variety of asteroida. D: Alai and Transalai Mts., central parts.

·          filipjevi Churkin, Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 71 – 72; figs. 3 b, 4 a, 6 d, 7 b, 9 c. TL: “Kazakhstan, North Tian-Shan, Zailiiskyi Alatau, Kaskelen R., 3300 - 3500 m”. Had been originally described as a subspecies of solona. D: North Tian Shan.

·          pletnevi Churkin, Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 66 – 69; figs. 2 a, 3 a, c, d, 4 e, 5 c, d, f, 6 e. TL: “Inn. Tian-Shan, Baydulu Mts., 5 km S Dolon Pass, 3000 – 3100 m”. Had been originally described as a subspecies of solona. D: Inner Tian Shan.

·          plyushchi Churkin, Kolesnichenko & Tuzov, 2000 (Melitaea). Helios, Moscow, 1: 69 – 71; figs. 2 b, 7 d, 9 d. TL: “Kirgizstan, Sary-Dzhas R., Kaingdy-Katta [sic!] Mts., 5 km W. Tashkoro v., 2900 – 3300 m”. Had been originally described as a subspecies of solona. D: known only from its type locality.

·          solona Alphéraky, 1881 (Melitaea). D: Dzhungarian Alatau Mts.

 

 

sultanensis Staudinger, 1886 (Melitaea). Stettiner entomologische Zeitschrift, 42: 234. TL: "Samarcand". Had been originally described as a variety of parthenie.

·          sultanensis Staudinger, 1886 (Melitaea). D: Gissar; Darvaz.

 

sutschana Staudinger, 1892 (Melitaea). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 6: 183. TL: "Sutschan". Had been originally described as a variety of didyma.

·          graeseri P.Gorbunov in Korshunov & Gorbunov, 1995 (Melitaea). Dnevnye babochki asiatskoj chasti Rossii, Ekaterinburg: 92. TL: "Сахалин, окр. г. Южносахалинска" [Sakhalin, Yuzhnosakhalinsk vicinity]. Had been originally described as a subspecies of sutschana. D: Sakhalin.

·          sutschana Staudinger, 1892 (Melitaea). D: Amur region; Ussuri region; South Transbaikal.

 

transcaucasica Turati, 1919 (Melitaea). Naturalisto Siciliense, 23: 225. TL: "Nucha". Had been originally described as a subspecies of didyma.

·          transcaucasica Turati, 1919 (Melitaea). D: Caucasus; Transcaucasus.

 

trivia [Denis & Schiffermüller], [1775] (Papilio). Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend, Wien: 179. TL: Vienna. Had been originally described as a good species.

·          caucasica Staudinger, 1861 (Melitaea). Catalog der Lepidopteren des europaeischen Faunengebietes (Aufl. 1), Berlin: 8. TL: "Kaukasus". Had been originally described as a variety of didyma. D: Caucasus; Transcaucasus.

·          nana Staudinger, 1871 (Melitaea). Catalog der Lepidopteren des europaeischen Faunengebietes (Aufl. 3), Berlin: 18. TL: "Türkei". Had been originally described as a variety of trivia. D: Kopet Dagh.

·          trivia [Denis & Schiffermüller], [1775] (Papilio). D: Central and southern part of ex-USSR's Europa; Caucasus; Kazakhstan; South Altai.

·          gulasinaria ssp.n. [nec singularia Korshunov in Korshunov  & Gorbunov, 1995, nomen nudum]

Diagnosis. From the nominotypical subspecies differs by more light and motley pattern and a little by extended form of wings.

Description. Labial palpi in yellow hairs. Length of a forewing is 18 mm. The wings from above light-red, motley, as black points are well allocated – fine, relief. From below on a back wing the black strokes thin, from the inside to red belt adjoin leaky. The average field light yellow, on colour does not differ from basal and external part. On an average field from forward edge up to the central cell two lines wrong black spots from each party – till four.

The description is translated from Korshunov, 1977: 668.

Material. Holotype female collected in the Khubsugul lake vicinity in North Mongolia, which deposite in the Institute of Ecology and Systematics of Animals in Novosibirsk.

D: Tuva.

 

turanica Erschoff in Fedtschenko, 1874 (Melitaea). Reise nach Turkestan, Sankt-Petersburg, 2 (5) (7): 47. TL: "Samarcand". Had been originally described as a good species.

·          pamira Staudinger, 1887 (Melitaea). Stettiner entomologische Zeitschrift, 48: 53. TL: "Transalai". Had been originally described as a good species. D: Darvaz; North-West Pamir; Transalai.

·          turanica Erschoff in Fedtschenko, 1874 (Melitaea). D: Gissar; West Tian Shan.

·          ssp? D: Kirghiz Mts.

 

turkmanica Higgins, 1940 (Melitaea). Entomologist, London, 73: 51. TL: "Askhabad". Had been originally described as a good species.

·          turkmanica Higgins, 1940 (Melitaea). D: West Kopet Dagh (Kara-Kala vicinity) (Devyatkin, 1987 a).

 

vedica Nekrutenko, 1975 (Melitaea). Journal of the Lepidopterist's Society, 29: 102 – 105; figs. 1 – 6. TL: "Armenia Mer Ved. Pr., Chosrov, 1300 – 1500 m". Had been originally described as a good species.

·          vedica Nekrutenko, 1975 (Melitaea). D: Transcaucasus.

 

uitasica Wagner, 1913 (Melitaea). Entomologische Mitteilungen, 2: 94. TL: "Uitas-Kette". Had been originally described as a variety of asteroida.

·          uitasica Wagner, 1913 (Melitaea). D: Dzhungaria; Terskey Ala-Too Mts.; Kungey Ala Too Mts.; Ketmen Mts.

 

uvarovi P.Gorbunov in Korshunov & Gorbunov, 1995 (Melitaea). Dnevnye babochki asiatskoj chasti Rossii, Ekaterinburg: 93. TL: "Оренбургская обл., с. Донское" [Orenburg reg., Donskoe]. Had been originally described as a good species.

·          uvarovi P.Gorbunov in Korshunov & Gorbunov, 1995 (Melitaea). D: South Ural.

 

Enumeratio Insectorum Musei Billbergii: 77. TS: Papilio athalia Rottemburg, 1775.

 

alatauica Staudinger, 1881 (Melitaea). Stettiner entomologische Zeitschrift, 42: 291. TL: Dzhungarian Alatau – by the lectotype (Higgins, 1955: 74). Had been originally described as a variety of parthenie.

·          alatauica Staudinger, 1881 (Melitaea). D: Dzhungaria.

 

ambigua Ménétriès in Schrenck, 1859 (Melitaea). Reisen und Forschungen im Amur-Lande, Sankt-Petersburg, 2: 24; pl. 2, fig. 4. TL: "Djai". Had been originally described as a variety of athalia.

·          ambigua Ménétriès in Schrenck, 1859 (Melitaea). D: Amur region.

·          kenteana Seitz, 1909 (Melitaea). Die Groß-Schmetterlinge der Erde. Die Groß-Schmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart, 1: 222; Taf. 67 A. TL: Kentei Mts. Had been originally described as a subspecies of athalia. D: South Siberian mountains.

·          mandschurica Fixsen, 1887 (Melitaea). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 302. TL: “Amurgebiete und aus Korea”. Had been originally described as a variety of athalia. D: Ussuri region.

·          sachalinensis Matsumura, 1925 (Melitaea). Journal of College of Agriculture in Sapporo, 15: 99. TL: "Nord-Saghalien (Rikowskoie)". Had been originally described as good species. D: Sakhalin.

 

athalia Rottemburg, 1775 (Papilio). Naturforscher, Halle, 6: 5. TL: Paris. Had been originally described as a good species.

·          asiae Verity, 1940 (Melitaea). Transactions of the Royal entomological Society of London, 91: 612. TL: "Sutschan". Had been originally described as a subspecies of athalia. D: Ussuri region.

·          athalia Rottemburg, 1775 (Papilio). D: Central part of ex-USSR's Europa; Siberia; Altai; Carpathians.

·          hyperborea Dubatolov, 1997 (Mellicta). Far Eastern Entomologist, Vladivostok, 44: 8, fig. 18 – 20. TL: Kamchatka. Had been originally described as a subspecies of athalia. D: Kamchatka.

·          lucifuga Fruhstorfer, 1916 (Melitaea). Archiv Naturgeschichten, 82 A (2): 4. TL: "Saratow". Had been originally described as a subspecies of athalia. D: South of European part; Caucasus; Transcaucasus; North-West Kazakhstan.

·          norvegica Aurivillius, 1888 (Melitaea). Nordens. Fjärilar.: 30. TL: "Dorefjeld", Norway. Had been originally described as a variety of athalia. D: North of European part.

 

aurelia Nickerl, 1850 (Melitaea). Synopsis Lepidopteres Fauna Böhmens: 12. TL: Breslau. Had been originally described as a good species.

·          aurelia Nickerl, 1850 (Melitaea). D: European part; Kazakhstan; West Siberia; Carpathians.

·          distans Higgins, 1955 (Mellicta). Transactions of the Royal entomological Society of London, 106: 80; pl. 1, 2, fig. 13. TL: "Thian-Shan, Tekkes Valley, approximately 43°N, 80°E". Had been originally described as a subspecies of aurelia. D: TL.

·          petricola Nekrutenko, 1978 (Melitaea). Doklady Akademii nauk Ukrainskoj SSR, Kiev, Seria B, 1978 (7): 647. TL: "Ай-Петринскаяяйла, Крым" [Aj-Petrinskaja Jajla, Crimea]. Had been originally described as a subspecies of aurelia. D: Crimea.

 

britomartis Assmann, 1847 (Melitaea). Entomologische Zeitschrift, Breslau, 1 (1): 2. TL: Breslau. Had been originally described as a good species.

·          amurensis Staudinger, 1892 (Melitaea). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 6: 186. TL: "Raddefskaja, Sutschan, Ussuri, Baikal". Had been originally described as a variety of aurelia. D: Siberia; Amur region.

·          britomartis Assmann, 1847 (Melitaea). D: European part; North Kazakhstan; Crimea.

·          latefascia Fixsen, 1883 (Melitaea). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 303. TL: "Korea". Had been originally described as a variety of parthenie. D: Ussuri region.

 

caucasogenita Verity, 1930 (Melitaea). [nom. nov. pro Melitaea athalia var. caucasica Staudinger, 1871, non Melitaea didyma var. caucasica Staudinger, 1861]. Entomological Record and Journal of Variation, London, 42:134. TL: “Armenia”. Had been originally established as a variety of athalia.

·          caucasogenita Verity, 1930 (Melitaea). D: Transcaucasus; Small Caucasus.

 

centralasiae Wnukowsky, 1929 (Melitaea). Zoologische Anzeiger, 83: 222 [nom. nov. pro Melitaea aurelia var. mongolica Staudinger, 1892, nec Melitaea maturna mongolica Staudinger, 1892]. TL: Kentei. Had been originally described as a variety of aurelia.

·          centralasiae Wnukowsky, 1929 (Melitaea). D: South Siberian mountains.

·          kolymskaya Higgins, 1955 (Mellicta). Transactions of the Royal entomological Society of London, 106: 84 – 85. TL: "Sredne Kolymsk, north-eastern Siberia, Province Yakutsk, 68° lat." Had been originally described as a modification of centralasiae. D: Far East.

 

menetriesi Caradja, 1895 (Melitaea). Deutsche entomologische Zeitschrift Iris, 8: 50 [nom. nov. pro Melitaea parthenie var. orientalis Ménétriès, 1859, nec Melitaea orientalis Herrich-Schäffer, 1843]. TL: Kamchatka and Amur ("Bouri"). Had been originally described as a variety of parthenie.

·          menetriesi Caradja, 1895 (Melitaea). D: Kamchatka.

 

plotina Bremer, 1861 (Melitaea). Bulletin de l’Academie des Sciences, Sankt-Petersburg, 3: 544. TL: Bureja Mts. and Ussuri. Had been originally described as a good species.

·          pacifica Verity, 1932 (Melitaea). Entomological Record, London, 44: 71. TL: "Transk-baical-Troizkossowsk. 800 m". Had been originally described as a subspecies of athalia. D: South Siberian mountains.

·          plotina Bremer, 1861 (Melitaea). D: Amur region; Ussuri region.

·          standeli Dubatolov, 1997 (Mellicta). Far Eastern Entomologist, 44: 6 – 8; fig. 16 – 17. TL: Salair. Had been originally described as a subspecies of plotina. D: Salair; Bugotak (both in West Siberia).

 

rebeli Wnukowsky, 1929 (Melitaea). Zoologische Anzeiger, 83: 222 [nom. nov. pro  Melitaea asteria var. altaica Staudinger, 1901, nec Melitaea didyma altaica Grum-Grshimailo, 1893]. TL: "Alt. sum. mont; (Tschuja mont.)". Had been originally described as a variety of asteria.

·          rebeli Wnukowsky, 1929 (Melitaea). D: Altai.

 

reticulata Higgins, 1955 (Mellicta). Transactions of the Royal entomological Society of London, 106: 21. TL: "Altai centr. mont." Had been originally described as a modification of athalia.

·         reticulata Higgins, 1955 (Mellicta). D: South Siberian mountains.

 

westsibirica Dubatolov in Korshunov, 1998 ([Mellicta]). Novye opisanija i utochnenija dlja knigi Dnevnye babochki asiatskoj chasti SSSR, Novosibirsk: 18. TL: “с. Новый Шарап Ордынского р-на Новосибирской обл.” [Novy Sharap in Ordynsk district of Novosibirsk region]. Had been originally described as a subspecies of menetriesi but without in combination with generic name.

·          westsibirica Dubatolov in Korshunov, 1998 ([Melitaea]). D: West Siberia.

Subfamily Argynninae Duponchel, [1835]

Magazin für Insektenkunde, 6: 283. TS: Papilio paphia Linnaeus, 1758.

 

adippe Rottemburg, 1775 (Papilio). Naturforscher, Halle, 6: 13. TL: Halle. Had been originally described as a good species.

·          adippe Rottemburg, 1775 (Papilio). D: European part; Crimea; Carpathians.

·          chalciope Jachontov, [1909] (Argynnis). Revue Russe d’Entomologie, Moscow, 8: 287. TL: Borzhom in Caucasus. Had been originally described as an aberration of adippe. D: Caucasus.

·          chrysodippe Staudinger, 1892 (Argynnis). Deutsche entomologische Zeitschrift Iris, 5: 333. TL: “Kentei”. Had been originally described as a variety of adippe. D: South Siberian mountains.

·          doii Matsumura, 1928 (Argynnis). Insecta Matsumurana, Sapporo, 2 (4): 191, 193. TL: Kuriles: “Shakotan, Tomari and Tobutsu”. Had been originally described as a subspecies of adippe. D: Kuril Islands; Sakhalin.

·          jainadeva Moore, 1864 (Argynnis). Entomological Monthly Magazine, 1: 135. TL: India. Had been originally described as a good species. D: West Pamir; Alai; Darvaz; Gissar; Kopet Dagh.

·          pallescens  Butler, 1873 (Argynnis). Cistula Entomologica, 1: 164. TL: Japan, Hokkaido, “Hakodadi”. Had been originally described as a good species. D: Ussuri region.

·          tianschanica Alphéraky, 1881 (Argynnis). Horae Societatis entomologicae Rossicae, Sankt-Petersburg, 16: 412. TL: "Tian-Chian, Jouldousse". Had been originally described as a variety of adippe. D: Dzhungaria; Tian Shan; South Siberian mountains; Tarbagatai; Saur.

·          xanthodippe Fixsen, 1887 (Argynnis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 263. TL: Corea. Had been originally described as a variety of adippe. D: Amur region.

 

aglaja Linnaeus, 1758  (Papilio). Systema Naturae... (ed. 10), 1: 481. TL: Sweden. Had been originally described as a good species.

·          aglaja Linnaeus, 1758 (Papilio). D: European part; Carpathians; Crimea; Kazakhstan.

·          auxo Jachontov, [1909] (Argynnis). Revue Russe d’Entomologie, Moscow, 8: 287. TL: “Бештау” [Beshtau] – vicinity of Pyatigorsk in North Caucasus. Had been originally described as a variety of aglaja. D: Caucasus; Transcaucasus.

·          fortuna Janson, 1877 (Argynnis). Cistula Entomologica, London, 2: 154. TL: Japan. Had been originally described as a good species. D: Kuriles; Sakhalin, Far East; Kamchatka; Siberia; Ussuri region; Amur region.

·          vitatha Moore, 1874 (Argynnis). Proceedings of the zoological Society of London, 1874: 368. TL: "Cashmere, N. side of Rajdingan Pass and Gurais". Had been originally described as a good species. D: Middle Asiatic mountains.

 

alexandra Ménétriès, 1832 (Argynnis). Catalogue raisonné des objets de zoologie recueillis dans un voyage au Caucase et jusqu’alors frontiиres actuelles de la Perse, enterpris par ordre de S.M.Empereur, Saint-Pétersbourg: 246. TL: “...de Zouvant, sur la chine de Talyche”. Had been originally described as a good species.

·          alexandra Ménétriès, 1832 (Argynnis). D: Transcaucasus; Kara-Kala vicinity.

 

anadiomene C. & R.Felder, [1862] (Argynnis). Wiener entomologischen Monatsschrift, 6: 25. TL: Central China. Had been originally described as a good species.

·          ella Bremer, 1864 (Argynnis). Mémoires de la Academie des Sciences, Saint-Pétersbourg, 8: 94; pl. 8, fig. 1. TL: Vladivostok. Had been originally described as a good species. D: Amur region; Ussuri region.

 

argyrospilata Kotzsch, 1938 (Argynnis). Entomologische Rundschau, 55 (31): 360. TL: “Chodja-Mahomed, Alpenwiessenzone, 2850 – 3000 m”. Had been originally described as a good species.

·               argyrospilata Kotzsch, 1938. D: West Pamir: Khorog vicinity.

 

hyperbius Linnaeus, 1767 (Papilio). Systema Naturae, ed. 12: 787. TL: China. Had been originally described as a good species.

·       hyperbius Linnaeus, 1767 (Papilio). D: Southern Ussuri region.

 

laodice Pallas, 1771 (Papilio). Reisen durch verschieden Provinzen des Russischen Reichs in den Jahren 1768 – 1774, Sankt-Petersburg, 1: 470. TL: South Russia, Volga region. Had been originally described as a good species.

·          laodice Pallas, 1771 (Papilio). D: Middle and South of European part; South Siberian mountains; South and Middle Ural.

·          ussurika Kardakov, 1928 (Argynnis). Entomologische Mitteilungen, Berlin, 17: 271. TL: Ussuri river. Had been originally described as a subspecies of laodice. D: Ussuri region; Amur region; Sakhalin; South Kuriles.

 

nerippe C. & R. Felder, [1862] (Argynnis). Wiener entomologischen Monatsschrift, 6: 24. TL: “Japonia”. Had been originally described as a good species.

·          coreana Butler, 1882 (Argynnis). Annales of the Magazin of the natural History, (5) 9: 15.  TL: “Posiette Bay”. Had been originally described as a good species. D: Ussuri region.

 

niobe Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1:  481. TL: Sweden. Had been originally described as a good species.

·          barkhatovi P.Y.Gorbunov, 2001 (Fabriciana). The butterflies of Russia: classification, genitalia, keys for identification, Ekatherinburg: 186. TL: “Россия, Алтай, п. Акташ” [Russia, Altai, Aktash]. Had been originally described as a subspecies of niobe. D: Central and South-Eastern Altai.

·          demavendis Gross & Ebert, 1975 (Argynnis). Journal of the Entomological Society of Iran, Teheran, Suppl. 1: 36. TL: Iran, Demavend Mt. Had been originally described as a subspecies of niobe. D: Talysh; South Transcaucasus.

·          gigantea Staudinger, 1871 (Argynnis). Catalog der Lepidopteren des europäischen Faunengebietes, Berlin: 21. TL: “Hyr.[cania]”. Had been originally described as a variety of niobe. D: Caucasus.

·          niobe Linnaeus, 1758 (Papilio). D: European part; Kazakhstan; Carpathians; Crimea; South Siberian mountains (without Central and South-Eastern Altai); Amur region; Ussuri region; Sakhalin.

·          orientalis Alphéraky, 1881 (Argynnis). Horae Societatis entomologicae Rossicae, Sankt-Petersburg, 16: 410. TL: "Tian-Chian". Had been originally described as a variety of niobe. D: North and Inner Tian Shan.

·          ornata Staudinger, 1901 (Argynnis). Catalog der Lepidopteren des palaearktischen Faunengebietes, Berlin (Aufl. 1): 16. TL: Fergana. Had been originally described as a variety of niobe. D: West Tian Shan; Gissar; Alai; Kopet Dagh.

·          shiva Wyatt & Omoto, 1966 (Argynnis). Entomops, Nicé, 5: 151; figs. TL: "monts Shiva, N.-E.Badachshan, 2,600 m". Had been originally described as a subspecies of niobe. D: Darvaz; Pamir.

 

pandora [Denis & Schiffermüller], [1775] (Papilio). Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend, Wien: 176. TL: Vienna. Had been originally described as a good species.

·          pandora [Denis & Schiffermüller], [1775] (Papilio). D: South of European part; Crimea; Kazakhstan; Caucasus; Transcaucasus.

·          pasargades Fruhstorfer (Argynnis). Internationale entomologische Zeitschrift, Berlin, 2 (12): 69. TL: “Alexandergebirge…, Nordpersien… und Ashabad”. Had been originally described as a subspecies of maja. D: Kopet Dagh; Middle Asiatic mountains; South Altai.

 

paphia Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 481. TL: Sweden. Had been originally described as a good species.

·          masandarensis Gross & Ebert, 1975 (Argynnis). Juornal of the Entomological Society of Iran, Teheran, Suppl. 1: 37. TL: Iran, Mazandaran. Had been originally described as a subspecies of paphia. D: Transcaucasus; Caucasus.

·          neopaphia Fruhstorfer, 1907 (Argynnis). Societas Entomologica, 22: 67. TL: “Amur”. Had been originally described as a variety of paphia. D: Amur region; Ussuri region; Sakhalin; Kuriles, Transbaikal; Kamchatka.

·          paphia Linnaeus, 1758 (Papilio). D: European part; Carpathians; Crimea; North Tian Shan; Dzhungaria; Altai; Sayan; Tuva; Praebaikal.

 

ruslana Motschulsky, 1866 (Argynnis). Bulletin de la Société impériale des Naturalistes de Moscou, 39: 117. TL: “...fleuve Amour depuis la Schilka jusqu’à Nikolaevsk”. Had been originally described as a good species.

·          ruslana Motschulsky, 1866 (Argynnis). D: Amur region; Ussuri region; Sakhalin.

 

vorax Butler, 1871 (Argynnis). Transactions of the Royal entomological Society of London, 1871: 403. TL: Shanghai in East China. Had been originally described as a good species.

·          vorax Butler, 1871 (Argynnis). D: Ussuri region.

 

zenobia Leech, 1890 (Argynnis). Entomologist, London, 23: 188. TL: “Ta-chien-lu”. Had been originally described as a good species.

·          penelope Staudinger, 1891 (Argynnis). Deutsche entomologische Zeitschrift Iris, 4: 339. TL: “Sutschan” [Partizansk in Ussuri region]. Had been originally described as a variety of zenobia. D: Southern Ussuri region.

 

Bulletin de la Société impériale des Naturalistes de Moscou, 24: 439. TS: Argynnis paulina Nordmann, 1851.

 

sagana Doubleday in Doubleday & Westwood, 1847 (Argynnis). Genera of diurnal Lepidoptera: pl. 21, fig. 1. TL: China. Had been originally described as a good species.

·          ilona Fruhstorfer, 1907 (Argynnis). Societas Entomologica, 22: 67. TL: Japan, Tsusima island. Had been originally described as a subspecies of sagana. D: Sakhalin.

·          nordmanni Korshunov, 1984 (Argynnis). Chlenistonogie i gelminty, Novosibirsk: 61; ris. 2 a. TL: "долинар. Зеи, районБлаговещенск – Москвитино" [Zea river valley, district of Blagovestshensk – Moskvitino]. Had been originally described as a subspecies of sagana. D: Amur region.

·          paulina Nordmann, 1851 (Argynnis). Bulletin de la Société impériale des Naturalistes de Moscou, 24: 439. TL: Kiachta. Had been originally described as a good species. D: Transbaikal.

·          relicta Korshunov, 1984 (Argynnis). Chlenistonogie i gelminty, Novosibirsk: 59 – 61; ris. 1, 2 b. TL: "ГорнаяШория, тайгавдолинеручья – притокар. Мрас-Сублизустья" [Gornaya Shoriya, the tayga in the valley of a stream – of a tributary of the Mras-Su river near the estuary]. Had been originally described as a subspecies of sagana. D: Altai; Sayan; Tuva; Praebaikal.

 

Verzeichniss bekannter Schmett[er]linge, Augsburg, TS: Papilio egista Cramer, [1780].

 

eugenia Eversmann, 1847 (Argynnis). Bulletin de la Société impériale des Naturalistes de Moscou, 20 (3): 68. TL: "in provincia Irkutskensi". Had been originally described as a good species.

·          eugenia Eversmann, 1847 (Argynnis). D: Siberia; Transbaikal; Far East; Kamchatka.

·          montana Bang-Haas, 1906 (Argynnis). Deutsche entomologische Zeitschrift Iris, 19: 127. TL: “Altai”. Had been originally described as a variety of eugenia. D: Altai; Sayan; Praebaikal; Tuva.

 

lathonia Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 481. TL: Sweden. Had been originally described as a good species.

·          lathonia Linnaeus, 1758 (Papilio). D: European part; Caucasus; Transcaucasus; Kazakhstan; Turkestan; Middle Asiatic mountains; Crimea; Carpathians; South Siberian mountains; Tarbagatai; Saur; Dzhungaria.

 

Verzeichniss bekannter Schmett[er]linge, Augsburg, (2): 30. TS: Papilio hecate [Denis & Schiffermüller], [1775].

 

daphne [Denis & Schiffermüller], [1775] (Papilio). Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend, Wien: 177. TL: Vienna. Had been originally described as a good species.

·          daphne [Denis & Schiffermüller], [1775] (Papilio). D: South of European part and Siberia; Kazakhstan; Crimea; Caucasus; Transcaucasus; Kopet Dagh.

·          fumidia Butler, 1882 (Argynnis). Annales of the Magazin of the Natural History, (5) 9: 16. TL: “Posiette Bay”. Had been originally described as a variety of daphne. D: South Siberian Mountains; Amur region; Ussuri region.

 

hecate [Denis & Schiffermüller], [1775] (Papilio). Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend, Wien: 179. TL: Vienna. Had been originally described as a good species.

·          alaica Staudinger, 1886 (Argynnis). Stettiner entomologische Zeitschrift, 47: 227. TL: “Alai”. Had been originally described as a variety of hecate. D: All Middle Asiatic mountains without Darvaz and Chatkal Mts.

·          hecate [Denis & Schiffermüller], [1775] (Papilio). D: South of European part; Carpathians; Crimea; Kazakhstan; South Siberian mountains.

·          laterica Tuzov, 1999 (Brenthis). Helios, Moscow, 1: 105. TL: “Tajikistan, Peter I Mts., Ganishou loc.”. Had been originally described as a subspecies of hecate. D: Darvaz.

·          tergemina Churkin & Zhdanko, 2002 (Brenthis). Helios, Moscow, 3: 131 – 133; pl. 10, figs. 13 – 15, 18. TL: “Kirgizstan, West Tian-Shan, Chatkal Range, Ravat-Sai River, 1600 – 1800 m”. Had been originally described as a subspecies of hecate. D: Chatkal Mts.

·          transcaucasica Wnukowsky, 1929 (Argynnis). Zoologische Anzeiger, 83: 222. [nom.nov. pro Argynnis hecate var. caucasica Staudinger, 1871, nec Argynnis  caucasica Lederer, 1852]. TL: Caucasus. Had been originally described as a variety of hecate.  D: Caucasus; Transcaucasus; Kopet Dagh.

 

ino Rottemburg, 1775 (Papilio). Naturforscher, Halle, 6: 19; Tab. 1, Fig. 3, 4. TL: Halle. Had been originally described as a good species.

·          amurensis Staudinger, 1887 (Argynnis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 147. TL: “Askold und Blagowestschensk”. Had been originally described as a variety of ino. D: Amur region; Ussuri region.

·          achasis Fruchstorfer, 1907 (Argynnis). Entomologische Zeitschrift, Frankfurt am Main, 21: 163. [nom. nov. pro Argynnis ino clara Staudinger, 1892, non Argynnis clara Blanchard, 1844]. TL: “Kentei”. Had been originally described as a variety of ino. D: Mountains around the Baikal Lake.

·          magadanica Kurentzov, 1970 (Argynnis). Bulavousye cheshuekrylye Dalnego Vostoka SSSR, LeningraD: 109; ros. 92. TL: "Магаданскаяобл." [Magadan reg.]. Had been originally described as a subspecies of ina. D: Chukotka; North of Magadan region.

·          paidicus Fruhstorfer, 1907 (Argynnis). Entomologische Zeitschrift, Frankfurt am Main, 21: 163. TL: “Altai”. Had been originally described as a variety of ino. D: Altai; Sayan; Tuva; Praebaikal.

·          parvimarginalis Nakahara, 1926 (Argynnis). Insecutor Inscit menstr., Washington, 14: 47. [nom. nov. pro Argynnis ino karafutonis Matsumura, 1925, non Argynnis thore karafutonis Matsumura, 1919]. TL: “Saghalien”. Had been originally described as a subspecies of ino. D: Sakhalin; Kuriles.

·          siopelus Fruhstorfer, 1907 (Argynnis). Entomologische Zeitschrift, Frankfurt am Main, 21: 163. TL: “Kamtschatka”. Had been originally described as a subspecies of ino.  D: Kamchatka.

·          trachalus Fruhstorfer, 1916 (Argynnis). Arkchiv Naturgeschichten, 82 (A): 14. TL: “Tien Schan”. Had been originally described as a subspecies of ino. D: Tian Shan.

·          vitimensis Wnukowsky, 1929 (Argynnis). Zoologische Anzeiger, 83: 223 [nom. nov. pro Argynnis ino borealis Seitz, 1908, nec Argynnis aglaja borealis Strand, 1901]. TL: Vitim. Had been originally described as a subspecies of ino. D: Far East.

 

mofidii Wyatt, 1968 (Brenthis). Zeitschrift der Wiener entomologischen Gesellschaft, Wien, 53: 29 – 30. TL: "Darband, north of Tehran in the Elburz range at 2250 m" (Leestmans & Carbonell, 1993: 123). Had been originally described as a subspecies of hecate.

·          mofidii Wyatt, 1968 (Brenthis). D: West Kopet Dagh.

·          zabensis Leestmans & Carbonell, 1993 (Brenthis). Linneana Belgica, Tervuren, 14 (3): 128 – 129; frontispiece; pl. 1, 5 – 8; fig. 2 B, 3 B. TL: "Dez Valley, 1700 – 2200 m, Hakkâri province". Had been originally described as a subspecies of mofidii. D: South Transcaucasus.

 

Lepidoptera Indica, London, 4: 243. TS: Papilio pales [Denis & Schiffermüller], [1775].

 

alaskensis Holland, 1900 (Brenthis). Entomological News, 11: 383. TL: "mountains between Forty-Mile and Mission Creeks". Had been originally described as a variety of pales.

·          alaskensis Holland, 1900 (Brenthis). D: North of Siberia and Far East; Kamchatka; Chukotka.

·          bato Churkin, 1999 (Boloria). Atalanta, Würzburg, 29: 117; pl. 9, fig. 4. TL: Northern Transbaikalia, Kodar Mts.: Sul’ban river. Had been originally described as a subspecies of alaskensis. D: Transbaikal.

·          sedykhi Crosson du Cormier, 1977 (Boloria). Alexanor, Paris, 10: 46. TL: "Oural polaire". Had been originally described as a subspecies of alaskensis. D: Polar Ural; Polar tundras of European part; Kola peninsula.

 

alberta Edwards, 1890 (Argynnis). Canadian Entomologist, 22: 113. TL: "Laggan, Alberta prov., Canada". Had been originally described as a good species.

·          kurenzovi Wyatt, 1961 (Clossiana). Entomologist’s Record and Journal of Variation, 73: 134. TL: Chukotka, mountains of Anyui river near Bilibino. Had been originally described as a subspecies of alberta. D: Chukotka: Providenija bay; Anyujsky Mts.

 

angarensis Erschoff, 1870 (Argynnis). Bulletin de la Société impériale des Naturalistes de Moscou, 43: 112. TL: "Gouvernement d’Irkoutsk”. Had been originally described as a good species.

·            alticola Sushkin & Tshetverikov, 1907 (Argynnis). Horae Societatis entomologicae Rossicae, Sankt-Petersburg, 38: 18 – 19. TL: "навысотахъуОйскагооз. иуАраданскагоперевала" [in the mountains near Ojskoje lake and Aradansky pass]. Had been originally described as a subspecies of angarensis. D: Altai; South Ural; West Siberia.

·            amurensis Staudinger, 1887 (Argynnis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 465. TL: Amur. Had been originally described as a variety of angarensis. D: Amur region; Sayan; Praebaikal; Tuva.

·          angarensis Erschoff, 1870 (Argynnis). D: Amur region; Transbaikal.

·          herzi Wnukowsky, 1926 (Argynnis). Mitteilungen der Münchener entomologischen Gesellschaft, 17: 70. TL: “Jakutsk”. Had been originally described as a subspecies of angarensis. D: Yakutia; Far East; Magadan region; Chukotka.

·          miyakei Matsumura, 1919 (Argynnis). Thous. Ins. Jap. Add., 3: 584; pl. 45, figs. 8, 9. TL: "Saghalien". Had been originally described as a subspecies of amathusia. D: Sakhalin; South Kuriles.

·          shelkovnikovi Korb & Yakovlev, 1997 (Boloria). Alexanor, Paris, 20: 45. TL: "Altaï, Oukok, Karaboulak, 2 200 – 2 500 m". Had been originally described as a subspecies of angarensis. D: Ukok plateau.

 

aquilonaris Stichel, 1908 (Argynnis). Berliner entomologische Zeitschrift, 53: 81; Taf. 3, fig. 5. TL: Sweden, Galliware. Had been originally described as a subspecies of pales.

·            aquilonaris Stichel, 1907 (Argynnis). D: North and area of European part.

·            huerteri Churkin & Bogdanov, 2003 (Boloria). Helios, Moscow, 4: 217 – 219; fig. 1, pl. 14, figs. 10 – 12. TL: “Russia, Siberia, Buryatia, Severobaikal’sk, Dawan Pass, 1400 m”. Had been originally described as a subspecies of aquilonaris. D: TL and West Sayan.

·            neopales Nakahara, 1926 (Argynnis). Insecutor Inscit. menstr., Washington, 14: 48. [nom. nov. pro Argynnis pales sachalinensis Matsumura, 1925, non Argynnis sachalinensis Matsumura, 1908]. TL: “North-Sakhalien (Rikovskoie)”. Had been originally described as a subspecies of pales. D: Sakhalin; South Kuriles.

·            roddi Kosterin, 2000 (Boloria). Far Eastern Entomologist, 86: 3. TL: Altai: Kosh-Agatch. Had been originally described as a subspecies of pales. D: Central and South-Eastern Altai.

·            sima  Churkin, 2000 (Boloria). Helios, Moscow, 1: 107 - 109. TL: “Russia, Moscow region, Taldoma distr., Rostovsky v.”. Had been originally described as a subspecies of aquilonaris. D: Central part of European Russia.

 

banghaasi Stichel in Seitz, [1908] (Argynnis). Die Groß-Schmetterlinge der Erde. Die Groß-Schmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart, 1: 230, Taf. 68, Abb. a. TL: “Kentei”. Had been originally described as a forma of pales.

·            banghaasi Stichel in Seitz, [1908] (Argynnis). D: South Siberia; Far East.

·            infans Churkin, 2000 (Boloria). Helios, Moscow, 1: 109 – 110. TL: “Chukotka, Bilibino, Bol’shoi Koperveem loc.”. Had been originally described as a subspecies of banghaasi. D: Chukotka.

 

caucasica Lederer, 1853 (Argynnis). Verhandlungen des zoologisch-botanischen Vereins, Wien, 2: 22 [nom. nov. pro  arsilache Herrich-Schäffer, [1851]]. TL: “am Kazbegi” – by the lectotype (van Oorschot & Wagener, 1990: 114). Had been originally described as a good species.

·          caucasica Lederer, 1853 (Argynnis). D: Caucasus.

·          petrovi  Churkin, 2002 (Boloria). Helios, Moscow, 3: 108 – 109; pl. 8, figs. 10 – 13. TL: “Russia, Daghestan, Caucasus Major, Rutul distr., west from Luchek v., 3000 – 3400 m, Charisina Mt.”. Had been originally described as a subspecies of caucasica. D: TL.

 

chariclea Schneider, 1794 (Papilio). Schneider Magazine, 5: 588. TL: Lapland. Had been originally described as a good species.

·          chariclea Schneider, 1794 (Papilio). D: Polar tundras of Europa and Siberia; Kamchatka.

·          tshuktsha Dubatolov & Korshunov, 1998 in Korshunov, 1998 (Clossiana). Novye opisanija i utochneniya dlya knigi Dnevnye babochki asiatskoj chasti Rossii, Novosibirsk: 21. TL: “100 кмЮПевека, В.Чукотка, Усть-Чаун” [100 km S Pevek, East Chukotka, Ust-Chaun]. Had been originally described as a subspecies of chariclea. D: Chukotka.

 

dia Linné, 1767 (Papilio). Systema Naturae... (ed. 12), 1: 785. TL: Austria. Had been originally described as a good species.

·          alpina Elwes, 1899 (Argynnis). Proceedins of the Zoological Society of London, 1899: 339. TL: Altai: “Ongodai”. Had been originally described as a variety of dia. D: Dzhungaria; Tarbagatai; Altai; Sayan; Saur; Tuva; Praebaikal.

·          dia Linné, 1767 (Papilio). D: European part (without north); Crimea; Ural; Siberia; Far East.

·          calida Jachontov, 1911 (Argynnis). Izvestija Kavkazskogo Museja, Tiflis, 5: 304. TL: Caucasus: Teberda. Had been originally described as a variety of dia. D: Caucasus; Transcaucasus.

·          semota Tuzov, 1999 (Clossiana). Helios, Moscow, 1: 105 - 106. TL: “Russia, Magadan Reg., Magadan vic.”. Had been originally described as a subspecies of dia. D: Magadan region.

 

distincta Gibson, 1920 (Brenthis). Reports of Canadian Arctic Expeditions 1913 – 1918, 3 (1) Part I (Lepidoptera): 25. TL: “Harrington Creek, Yukon territory, Canada: lat. 65º05´”. Had been originally described as a good species.

·          dubatolovi Korshunov, 1987 (Clossiana). Nasekomye, klestshi i gelminty, Novosibirsk: 11 – 12; ris. 3. TL: “Хамар-Дабан, 15 кмюжнеепос. Слюдянка, вершинапикаЧерского, 2090 мнадур.м., уосыпи” [Khamar-Daban, 15 km S Sljudanka, the summit of the Cherski peak, 2090 m above sea level, near strew]. Had been originally described as a subspecies of distincta. D: Transbaikal.

·          ershovi Korshunov & Gorbunov, 1995 (Clossiana). Dnevnye babochki asiatskoj chasti Rossii, Ekaterinburg: 108. TL: “п. Хаптагайнар. Лена” [Khaptagai on the Lena river]. Had been originally described as a subspecies of astrate. D: Lena plateau in Yakutia.

·          machati Korshunov, 1987 (Clossiana). Nasekomye, klestshi i gelminty, Novosibirsk: 10 – 11; ris. 2. TL: “ПолярныйУрал, 112-йкилометржелезнойдорогиСейда – Лабытнанги” [Polar Ural, km 112 of the railway Seida – Labytnangi]. Had been originally described as a subspecies of distincta. D: North-East of European part; Polar Ural; north-west of Siberia.

·          tschukotkensis Wyatt, 1961 (Clossiana). Entomologist’s Record and Journal of Variation, 73: 134. TL: Chukotka, mountains on Anyui river near Bilibino. Had been originally described as a subspecies of distincta. D: Chukotka.

 

dulkeiti Kurentzov, 1970 (Brenthis). Bulavousye cheshuekrylye Dalnego Vostoka SSSR, LeningraD: 83, 108 – 109; ris. 76 (3). TL: “в альпийской зоне Колымского хребта (Омсукчанские горы, Мадаунские гольцы)” [in the alpine zone of Kolimsky Mts (Omsukchanskie mountains, Madaunskie goltsy)]. Had been originally described as a good species.

·          dulkeiti Kurentzov, 1970 (Brenthis). D: Far East (northern part).

 

erda Christoph, 1893 (Argynnis). Deutsche entomologische Zeitschrift Iris, 6: 87. TL: “Vilui”. Had been originally described as a good species.

·          erda Christoph, 1893 (Argynnis). D: Chukotka; north of Far East.

·          kitoica Belik, 1996 (Clossiana). Phegea, Antverpen, 24: 163 – 166; fig. 6; pl. 2, figs. 1 – 6. TL: “Russia, Vostochnyy Sayan mountains, khrebet (mountain range) Kitoiskiye Gol’tsy, river Kitoi upper stream, vicinity of the lake Il’chir, 2350 m”. Had been originally described as a subspecies of erda. D: TL.

 

erubescens Staudinger, 1901 (Argynnis). Catalog der Lepidopteren des palaearktischen Faunengebietes, Berlin (Aufl. 1): 35. TL: “Korla”. Had been originally described as a variety of hegemone.

Remark. “Die Art wurde zum erstenmal unter dem Namen Argynnis hegemone (Staudinger, 1881,  Stett. Ent. Ztg., 42: 292) beschrieben. Als Typenfundort von hegemone wurden von Staudinger “Ala Tau”, “Margelan” und “Tianschan” genannt und von Wagner, 1913 auf “Margelan” eingeschränkt. Dieser Name ist aber ungültig, da er ein Homonym von Argynnis hegemone (Godart, 1819, Ency. Meth., 9 (1): 238) ist” (Lukhtanov & Lukhtanov, 1994: 195).

·       ali Churkin, 2002 (Clossiana). Helios, Moscow, 3: 162 – 164; figs. 1 e, f, 2 c, f, pl. 12, figs. 13 – 18. TL: “Alai (northern slopes), Kollektorsky Mts., Dugoba R.”. Had been originally described as a subspecies of hegemone. D: North Alai.

·       ancilla Churkin, 2004 (Clossiana). Helios, Moscow, 5: 132 – 133; pl. 9, figs. 9 – 12. TL: “Tadjikistan, Peter the Great Range, 23 km SE Tadzhikobad [sic!], Ganishou v.”. Had been originally described as a subspecies of hegemone. D: Darvaz.

·       chotana Bang-Haas, 1915 (Argynnis). Deutsche entomologische Zeitschrift Iris, 29: 100. TL: Chotan in China. Had been originally described as a variety of hegemone. D: West Pamir.

·       erubescens Staudinger, 1901 (Argynnis). D: Alai; Gissar.

·       eximia Churkin, 2002 (Clossiana). Helios, Moscow, 3: 164 – 165; figs. 1 a, b, 2 a, d, pl. 12, figs. 1 – 6. TL: “Fergansky Mts., 15 km E Kalmak-Krchin v., Kugart R., 2000 – 2200 m”. Had been originally described as a subspecies of hegemone. D: Fergansky Mts.

·       haberhaueri Hemming, 1933 (Boloria). Entomologist, London, 66: 275 [nom. nov. pro Argynnis hegemone Staudinger, 1881, nec Argynnis hegemone Godart, 1819]. TL: “Margelan”. Had been originally described as a good species. D: North Tian Shan without Kirghiz Mts.

·       panterai Korb 2005. D: Kirghiz Mts.

·       tienschanica Wagner, 1913 (Argynnis). Entomologische Mitteilungen, 2: 114. TL: “Uitas-Kette”, “Juldus”. Had been originally described as a subspecies of hegemone. D: Inner Tian Shan.

·       tuzovi Churkin, 2002 (Clossiana). Helios, Moscow, 3: 165 – 167; figs. 1 c, d, 2 b, e, pl. 12, figs. 7 – 12. TL: “Talassky Alatau, Alabel Pass, 3300 m”. Had been originally described as a subspecies of hegemone. D: West Tian-Shan.

 

eunomia Esper, [1799] (Papilio). Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen, Erlangen, 1: Tab. 110, Fig. 5. TL: “Königsberg”. Had been originally described as a good species.

·       acidalia Böber, 1809 (Papilio). Mémoires de la Société des naturalistes de Moscou, 2: 306. TL: “de la Sibérie”. Had been originally described as a good species. D: Mountains of South Siberia.

·       asiatica Staudinger, 1901 (Argynnis). Catalog der Lepidopteren des palaearktischen Faunengebietes, Berlin (Aufl. 1): 34. TL: “Sib.[eria] m.[eridionalis] (Alt[ai] Saj[an]; Kent.[ei]); Amur”. Had been originally described as a variety of aphirape. D: South and middle Siberia; Amur region; Ussuri region; Sakhalin.

·       eunomia Esper, [1799] (Papilio). D: Central area of European part; Caucasus.

·       ossianus Herbst, 1800 (Papilio). Natursystematic der Insekten. Schmetterlinge, 10: 98, 270; Fig. 4, 5. TL: [North Europa]. Had been originally described as a good species. D: North of Euroasia; Far East; Kamchatka.

 

euphrosyne Linnaeus, 1758 (Papilio). Systema Naturae... (ed. 10), 1: 481. TL: Sweden. Had been originally described as a good species.

·       anka Fruhstorfer, 1907 (Argynnis). Entomologische Zeitschrift, Frankfurt am Main, 21: 164. TL: “Kaukasus”. Had been originally described as a subspecies of euphrosyne. D: Caucasus; Transcaucasus.

·       euphrosyne Linnaeus, 1758 (Papilio). D: North and Middle of European part and Siberia.

·       kamtschadalus Stichel in Seitz, [1908] (Argynnis). Die Groß-Schmetterlinge der Erde. Die Groß-Schmetterlinge des palaearktischen Faunengebietes. Die palaearktischen Tagfalter, Stuttgart, 1: 233. TL: “Kamtschatka”. Had been originally described as a subspecies of euphrosyne. D: Kamchatka.

·       rusalka Fruhstorfer, 1909 (Argynnis). Internationale entomologische Zeitschrift, Guben, 3: 113. TL: “Saratow”. Had been originally described as a subspecies of euphrosyne. D: South of European part; North Kazakhstan.

·       orphanoides Huang & Murayama, 1992 (Clossiana). Tyô to Ga, 43: 7; fig. 17. TL: “Jiadengyu, Altai, 1380 – 1600 m, grassland in forest zone”. Had been originally described as a subspecies of euphrosyne. D: SW Altai: Ukok plateau.

·       ssp? D: Kirghiz Mts.: Chon-Kuurchak valley.

 

franciscana Verhulst, 1999 (Clossiana). Lambillionea, Bruxelles, 99 (2): 213 – 214; textfigs. 1 – 4; col. pl. figs. 1 – 9. TL: «Pakistan, Gilgit, col de Kunjerab 4000 – 4200m». Had been originally described as a good species.

·       franciscana Verhulst, 1999 (Clossiana) (an bona ssp?). D: «Clossiana franciscana vole aussi au Kirgizistan; j’ai en effet capturé un mâle dans l’Alaï aux environs du village de Madjan 2500 – 3000m le 21/7/1996. P.François en a également pris deux le 23/7/1996 dans la même localité!» (Verhulst, 1999: 213).

 

freja Thunberg & Becklin, 1791 (Papilio). Dissertatio entomologicae sistens Insecta Svecica, 2: 34; Tab. 5, fig. 14. TL: [Sweden]. Had been originally described as a good species.

·       freja Thunberg & Becklin, 1791 (Papilio). D: North of European part; Central Siberia; Transbaikal; Kamchatka; Amur region; Yakutia.

·       pallida Elwes, 1899 (Argynnis). Transactions of the Royal entomological Society of London, 1899: 338. TL: “south of Kosh-Agach, at 7000 – 8000 feet”. Had been originally described as a variety of freja. D: Altai; Sajan; Tuva (north); Praebaikal.

·       zamolodchikovi Churkin, 2001 (Clossiana). Helios, Moscow, 2: 99 – 101; pl. 8, figs. 10 - 12. TL: “Russia, East Chukotka, 13 km SE Ioni L.”. Had been originally described as a subspecies of freija [sic!]. D: Chukotka.

 

frigga Thunberg & Becklin, 1791 (Papilio). Dissertatio entomologicae sistens Insecta Svecica, 2: 33. TL: Lapland. Had been originally described as a good species.

·       alpestris Elwes, 1899 (Argynnis). Transactions of the Royal entomological Society of London, 1899: 340. TL: “high Tchuja Mts. at from 7000 – 8000 feet elevation”. Had been originally described as a variety of frigga. D: Altai; Sayan; Praebaikal; Tuva.

·       annae Suschkin, 1906 (Argynnis). Revue Russe d’Entomologie, Moscow, 6: 5. TL: “die alpine Region der Kette Saur am Berge Musstau”, “zwischen Uidene und Tschagan-Obo”. Had been originally described as a subspecies of frigga. D: Tarbagatai Mts. in South Altai.

·       frigga Thunberg & Becklin, 1791 (Papilio). D: Polar tundra of European part and Siberia; Chukotka; Kamchatka.

·       maritima Kardakov, 1928 (Argynnis). Entomologische Mitteilungen, Berlin, 17: 270, Taf. 7, Abb. 4. TL: Terney Bay. Had been originally described as a variety of frigga. D: Amur region; Sykhote Alin.

·       ssp? D: Sakhalin: “Окрестность, ЗаливПильтун, ОхинскийРайон, Сахалин” [Piltun bay vicinity, Okhinsk district, Sakhalin] (Takahashi, 1994: 59).

 

frigidalis Warren, 1944 (Boloria). Transactions of the Royal entomological Society of London, 94: 48, pl. 33, figs. 184-185. TL: “Tchuja Mountains”, “Arasan”. Had been originally described as a forma of napaea.

·       frigidalis Warren, 1944 (Boloria). D: Altai.

·       purpurea Churkin, 1999 (Boloria). Atalanta, Würzburg, 29: 118; pl. 9, figs. 1 - 3. TL: “Russia, Buryatia, Barguzin Mts., Nesterikha River, Kedrovoe Lake, at foot of Haramook Peak”. Had been originally described as a good species. D: Barguzin Mts.

 

generator Staudinger, 1886 (Argynnis). Stettiner entomologische Zeitschrift, 47: 235. TL: “Alai (Margelan), Osch, Usgent, Namangan, Tianschan, Dshungarischer Alatau”. Had been originally described as a variety of pales.

·       generator Staudinger, 1886 (Argynnis). D: Tian Shan; Dzhungaria; Saur; Tarbagatai; Alai.

 

hakutozana Matsumura, 1927 (Argynnis). Insecta Matsumurasca, 1: 165. TL: “Hakuto”. Had been originally described as a good species.

·       hakutozana Matsumura, 1927 (Argynnis). D: South of Ussuri region.

 

iphigenia Graeser, 1888. Berliner entomologische Zeitschrift, 32: 90. TL: “Nicol.[aefsk]”. Had been originally described as a good species.

·       iphigenia Graeser, 1888. D: Ussuri region; Amur region; South Sikhote Alin.

·       sachalinensis Matsumura, 1925 (Argynnis). Entomologische Zeitschrift, Stuttgart, 20: 160. TL: Sakhalin: Oguma. Had been originally described as a good species. D: Sakhalin; South Kuriles.

 

improba Butler, 1877 (Argynnis). Entomological Montley Magazine, 13: 206. TL: “Winter Cove and Cambridge Bay”. Had been originally described as a good species.

·       improbula Bryk, 1920 (Argynnis). Entomologische Tidskrift, Leiden, 41: 131, figs. 2, 5. TL: “Tornetrask”. Had been originally described as a forma of improba. D: zone of tundras of Eurasia.

 

matveevi P.Gorbunov & Korshunov, 1995 (Clossiana). Dnevnye babochki asiatskoj chasti Rossii, Ekaterinburg: 109. TL: “Алтай, пос. Акташ, 2700 м”. Had been originally described as a good species.

·       matveevi P.Gorbunov & Korshunov, 1995 (Clossiana). D: TL.

 

napaea Hoffmannsegg, 1804 (Papilio). Magazin für Insektenkunde, 3: 196. TL: Alps of Tyrol. Had been originally described as a good species.

·       altaica Grum-Grshimailo, 1893 (Argynnis). Horae Societatis Entomologicae Rossicae, Sankt-Petersburg, 27: 128. TL: “In Montibus Altaicis (fluv. Kara-Kaba)”. Had been originally described as a variety of pales. D: Altai; Sayan; Tuva; Praebaikal.

·       napaea Hoffmannsegg, 1804 (Papilio). D: North of European part and Siberia.

·       pustagi Korshunov & Ivonin in Korshunov & Gorbunov, 1995 (Boloria). Dnevnye babochki asiatskoj chasti Rossii, Ekaterinburg: 109. TL: “ГорнаяШория, окр. п. Шерегеш, г. Пустаг, 1000 – 1300 м” [Gornaya Shoriya, Sheregesh vicinity, Pustag Mt., 1000 – 1300 m]. Had been originally described as a subspecies of altaica. D: Kusnezky Alatau Mts.

·       vinokurovi Dubatolov, 1992 (Boloria). Vestnik zoologii, Kiev, 1992 (6): 40 – 41. TL: “Якутия, Верхоянскийхр., р. Нямни, лев. истокр. Кокчин” [Yakutia, Verkhoyansky Mts., Nyamni river, the left source of Kokchin river]. Had been originally described as a subspecies of napaea. D: Far East.

 

natazhati Gibson, 1920 (Argynnis). Report on the Canadian arctic expedition 1913-18. 3: 211 – 221. TL: «Mt Natazhat, St Elias Mts., Yukon Territory». Had been originally described as a good species.

·       perunovi Korb, 1999 (Clossiana). Lambillionea, Bruxelles, 99 (4): 570; fig. 1. TL: «Russia, Chita reg., Udokan, 1000m». Had been originally described as a subspecies of natazhati. D: TL.

 

oscarus Eversmann, 1844 (Argynnis). Bulletin de la Société des Naturalistes de Moscou, 17: 588; Tab. 14, fig. 1. TL: “in provincia Irkutzkensi”. Had been originally described as a good species.

·       australis Graeser, 1888 (Argynnis). Berliner entomologische Zeitschrift, 32: 90. TL: “Wlad.[iwostok]”. Had been originally described as a variety of oscarus. D: Amur region; Ussuri region.

·       insularia Korshunov, 1996 (Clossiana). Dopolnenija i ispravlenija k knige Dnevnye babochki asiatskoj chasti Rossii, Novosibirsk: 28 [nom. nov. pro sachalinensis Matsumura, 1936, nec sachalinensis Matsumura, 1925]. TL: “Saghalien”. Had benn originally described as a subspecies of oscarus. D: Sakhalin; South Kuriles.

·       oscaroides Ménétriès, 1859 (Argynnis). Bull. phys. Acad. St.-Pétersb., 3 (1): 216. TL: [Okhotsk]. Had been originally described as a variety of oscarus. D: Lena plateau, mountains of South-Eastern Siberia.

·       oscarus Eversmann, 1844 (Argynnis). D: South Siberia; South Ural; Middle Ural; midddle area of East European part; Transbaikal.

 

perryi Butler, 1882 (Argynnis). Annales of the Magazin of the natural History, (5) 9: 16. TL: “Pasiette Bay, N.E. Corea”. Had been originally described as a good species.

·       perryi Butler, 1882 (Argynnis). D: Ussuri region.

 

polaris Boisduval, 1829 (Argynnis). European Lepidoptera. Index methodologique, Paris: 15. TL: “Cap Nord”. Had been originally described as a good species.

·       digna Churkin, 2001 (Clossiana). Helios, Moscow, 2: 97 – 99; pl 8, figs. 4 – 6. TL: “Russia, Wrangel Is., Neizvestnaya R. (middle stream)”. Had been originally described as a subspecies of polaris. D: Wrangel Island.

·       polaris Boisduval, 1829 (Argynnis). D: Polar tundra of Euroasia; Chukotka; north of Magadan region.

 

selene [Denis & Schiffermüller], [1775] (Papilio). Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend, Wien: 321. TL: Vienna. Had been originally described as a good species.

·       dilutior Fixsen, 1887 (Argynnis). Mémoires sur les Lépidoptères, Saint-Pétersbourg, 3: 167. TL: Pung-Tung in Corea. Had been originally described as a variety of selene. D: Amur region; Ussuri region; Sakhalin; Kuriles.

·       selene [Denis & Schiffermüller], [1775] (Papilio). D: European part; Caucasus; North Kazakhstan; Siberia.

 

selenis Evermann, 1837 (Argynnis). Bulletin de la Société des Naturalistes de Moscou, 10: 10. TL: “Kasanischer Gouvernement”. Had been originally described as a good species.

·       kononovi Kurentzov, 1970 (Brenthis). Bulavousye cheshuekrylye Dalnego Vostoka SSSR, LeningraD: 102. TL: “Магаданскаяобл.: Мадаунскиегольцы” [Magadan region: Madaunskye goltsy] – by the syntypes (Azarova, 1986: 125). Had been originally described as a subspecies of selenis. D: North of Far East; Magadan region; Chukotka.

·       selenis Eversmann, 1837 (Argynnis). D: European part; North Kazakhstan; ? Caucasus.

·       sibirica Erschoff, 1870 (Argynnis). Bulletin de la Societe imperiale des Naturalistes de Moscou, 43: 112. TL: “Altai et Sibérie orientale”. Had been originally described as a variety of selenis. D: Siberia; south of Far East; Amur region; Ussuri region; Sakhalin.

 

speranda Grosser, 1979 (Clossiana). Reichenbachia, Leipzig, 17: 331 – 332; Abbn. 1, 2, 3 b. TL: “Ulangom / Charchira, NW-Mongolei”. Had been originally described as a good species.

·       speranda Grosser, 1979 (Clossiana). D: Tuva.

 

thore Hübner, [1803] (Papilio). Sammlung europäischer Schmetterlinge, Augsburg, 1: Fig. 571 – 573. TL: “Tiroler Alpen”. Had been originally described as a good species.

·       borealis Staudinger, 1861 (Argynnis). Catalog der Lepidopteren des europaeischen Faunengebietes, Berlin (Aufl. 1): 9. TL: Lapland. Had been originally described as a variety of thore. D: North of Europe.

·       hyperusia Fruhstorfer, 1907 (Argynnis). Entomologische Zeitschrift, Frankfurt am Main, 21: 165. TL: “Amur, …Sibirien”. Had been originally described as a subspecies of thore. D: Far East; Kamchatka; Transbaikal; Sakhalin; Amur region; Ussuri region.

·       lacrimosa Korb, 1999 (Clossiana). Lambillionea, Bruxelles, 99 (4): 571; fig. 3. TL: «Altai, Ukok, Kara-Bulak». Had been originally described as a subspecies of thore. D: South Altai.

·       ssp? D: Central area of the European part.

 

titania Esper, [1793] (Papilio). Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen, Erlangen, 1: 58. TL: “Sardinien”. Had been originally described as a good species.

·       bivina Fruhstorfer, 1908 (Argynnis). Internationale entomologische Zeitschrift, Guben, 1: 310. TL: “Saratow”. Had been originally described as a subspecies of amathusia. D: European part; Ural; West Siberia.

·       staudingeri Wnukowsky, 1929 (Brenthis). Zoologische Anzeiger, 83: 222 [nom. nov. pro Argynnis amathusia var. sibirica Staudinger, 1892, nec Argynnis selenis var. sibirica Erschoff, 1870]. TL: “Kentei”. Had been originally described as a variety of amathusia. D: South Siberia; Transbaikal; Amur region; Ussuri region; Sakhalin.

 

tritonia Boeber, 1812 (Papilio). Mémoires de la Société impériale des Naturalistes de Moscou, 3: 20; pl. 1, fig. 1, 2. TL: “Sibérie”. Had been originally described as a good species.

·       elatus Staudinger, 1892 (Argynnis). Deutsche entomologische Zeitschrift Iris, 5: 329. TL: “Kentei”. Had been originally described as a variety of amphilochus. D: South Siberian mountains; Amur region.

·       tritonia Boeber, 1812 (Papilio). D: Middle Siberia; Ussuri region; Middle Far East.